Jose Negron, PhD
Research Entomologist
jose.negron@usda.gov
Treesearch Publications
- Andrew M. Liebhold, Matthew W. Ethington, Brett Stadsvold, Jose Negron. 2025. Spread of the emerald ash borer, Agrilus planipennis, in the Front Range region of Colorado
- Paula J. Fornwalt, Andreas P. Wion, Anna W. Schoettle, Miranda D. Redmond, Steven Alton, Javier E. Mercado, Brice Hanberry, Jose Negron. 2024. Vulnerability of major Colorado Front Range tree species to climate change (Chapter 5)
- Jose Negron, Javier E. Mercado. 2024. Insects and diseases and climate change in Colorado Front Range forests (Chapter 8)
- Nehalem C. Clark, Signe Leirfallom, Hannah Farrell, Michael A. Battaglia, Brandon M. Collins, Justin S. Crotteau, Kimberley T. Davis, Paula J. Fornwalt, Paul F. Hessburg, Sharon M. Hood, Eric E. Knapp, W. Keith Moser, Jose Negron. 2024. What do we know about forest treatments and fire? Photos and long-term studies help us understand
- Kyle C. Rodman, Robert A. Andrus, Amanda R. Carlson, Trevor A. Carter, Teresa B. Chapman, Jonathan D. Coop, Paula J. Fornwalt, Nathan S. Gill, Brian J. Harvey, Ashley E. Hoffman, Katharine C. Kelsey, Dominik Kulakowski, Daniel C. Laughlin, Jenna E. Morris, Jose Negron, Katherine Nigro, Gregory S. Pappas, Miranda D. Redmond, Charles C. Rhoades, Monique E. Rocca, Zoe H. Schapira, Jason S. Sibold, Camille S. Stevens-Rumann, Thomas T. Veblen, Jianmin Wang, Xiaoyang Zhang, Sarah J. Hart. 2022. Rocky Mountain forests are poised to recover following bark beetle outbreaks but with altered composition
- Nancy Sturdevant, Laurel Haavik, Jose Negron. 2022. Douglas-fir Tree Mortality Caused by the Douglas-fir Beetle in Thinned and Unthinned Stands in Montana, USA
- Amanda R. Carlson, Jason S. Sibold, Jose Negron. 2021. Wildfire and spruce beetle outbreak have mixed effects on below-canopy temperatures in a Rocky Mountain subalpine forest
- Brian J. Harvey, Robert A. Andrus, Michael A. Battaglia, Jose Negron, Alexandra Orrego, Thomas T. Veblen. 2021. Droughty times in mesic places: Factors associated with forest mortality vary by scale in a temperate subalpine region
- Jose Negron, Dan W. McCollum. 2020. Taking the long view: Mountain pine beetles as agents of change
- Jose Negron, Robert Pate, Justin D. Derner. 2020. Flight of the Mountain Pine Beetle, Dendroctonus ponderosae Hopkins (Coleoptera: Curculionidae: Scolytinae), in Suburban Cheyenne, Wyoming, USA during Summer 2011
- Justin B. Runyon, Christopher J. Fettig, Jared Trilling, A. Steven Munson, Leif A. Mortenson, Brytten E. Steed, Kenneth E. Gibson, Carl L. Jørgensen, Stephen Mckelvey, Joel D. McMillin, Jackson Audley, Jose Negron. 2020. Changes in understory vegetation including invasive weeds following mountain pine beetle outbreaks
- Amanda R. Carlson, Jason S. Sibold, Jose Negron. 2020. Canopy structure and below-canopy temperatures interact to shape seedling response to disturbance in a Rocky Mountain subalpine forest
- Jose Negron. 2020. Within-stand distribution of tree mortality caused by mountain pine beetle, Dendroctonus ponderosae Hopkins
- M. M. Dudley, N. A. Tisserat, W. R. Jacobi, Jose Negron, J. E. Stewart. 2020. Pathogenicity and distribution of two species of Cytospora on Populus tremuloides in portions of the Rocky Mountains and midwest in the United States
- Jackson P. Audley, Christopher J. Fettig, A. Steven Munson, Justin B. Runyon, Leif Mortenson, Brytten E. Steed, Kenneth E. Gibson, Carl L. Jørgensen, Stephen Mckelvey, Joel D. McMillin, Jose Negron. 2020. Impacts of mountain pine beetle outbreaks on lodgepole pine forests in the Intermountain West, U.S., 2004–2019
- Jose Negron, Laurie Huckaby. 2020. Reconstructing historical outbreaks of mountain pine beetle in lodgepole pine forests in the Colorado Front Range
- Osiris Valerio-Mendoza, Jazmin Garcia-Roman, Moises Becerril, Francisco Armendariz-Toledano, Gerardo Cuellar-Rodriguez, Jose Negron, Brian Sullivan, Gerardo Zuniga. 2019. Cryptic species discrimination in western pine beetle, Dendroctonus brevicomis LeConte (Curculionidae: Scolytinae), based on morphological characters and geometric morphometrics
- Jose Negron. 2019. Biological aspects of mountain pine beetle in lodgepole pine stands of different densities in Colorado, USA
- Jose Negron, Robert Cain, Andy Cadenhead, Brian Waugh. 2019. A test of lethal trap trees for control of spruce beetles
- Adrianna C. Foster, Jacquelyn K. Shuman, Herman H. Shugart, Jose Negron. 2018. Modeling the interactive effects of spruce beetle infestation and climate on subalpine vegetation
- Jose Negron, Bob Cain. 2018. Mountain pine beetle in Colorado: A story of changing forests
- Howard Williams, Sharon M. Hood, Christopher R. Keyes, Joel M. Egan, Jose Negron. 2018. Subwatershed-level lodgepole pine attributes associated with a mountain pine beetle outbreak
- Danielle M. Malesky, Barbara J. Bentz, Gary R. Brown, Andrea R. Brunelle, John M. Buffington, Linda M. Chappell, R. Justin DeRose, John C. Guyon, Carl L. Jorgensen, Rachel A. Loehman, Laura L. Lowrey, Ann M. Lynch, Marek Matyjasik, Joel D. McMillin, Javier E. Mercado, Jesse L. Morris, Jose Negron, Wayne G. Padgett, Robert Progar, Carol B. Randall. 2018. Effects of climate change on ecological disturbances [Chapter 8]
- Jose Negron, Jennifer G. Klutsch. 2017. Probability of infestation and extent of mortality models for mountain pine beetle in lodgepole pine forests in Colorado
- Amanda R. Carlson, Jason S. Sibold, Timothy J. Assal, Jose Negron. 2017. Evidence of compounded disturbance effects on vegetation recovery following high-severity wildfire and spruce beetle outbreak
- O. Valerio-Mendoza, F. Armendariz-Toledano, G. Cuellar-Rodriguez, Jose Negron, G. Zuniga. 2017. The current status of the distribution range of the western pine beetle, Dendroctonus brevicomis (Curculionidae: Solytinae) in northern Mexico
- Jose Negron, Kurt K. Allen, Angie Ambourn, Blaine Cook, Kenneth Marchand. 2017. Large-scale thinning, ponderosa pine, and mountain pine beetle in the Black Hills, USA
- Jose Negron, John B. Popp. 2017. Can spruce beetle (Dendroctonus rufipennis Kirky) pheromone trap catches or stand conditions predict Engelmann spruce (Picea engelmannii Parry ex Engelm.) tree mortality in Colorado?
- Adrianna C. Foster, Jacquelyn K. Shuman, Herman H. Shugart, Kathleen A. Dwire, Paula J. Fornwalt, Jason Sibold, Jose Negron. 2017. Validation and application of a forest gap model to the southern Rocky Mountains
- Adrianna C. Foster, Jonathan A. Walter, Herman H. Shugart, Jason Sibold, Jose Negron. 2017. Spectral evidence of early-stage spruce beetle infestation in Engelmann spruce
- Daniel R. West, Jennifer S. Briggs, William R. Jacobi, Jose Negron. 2016. Mountain pine beetle host selection between lodgepole and ponderosa pines in the southern Rocky Mountains
- Jonathan A. Cale, Jennifer G. Klutsch, Nadir Erbilgin, Jose Negron, John D. Castello. 2016. Using structural sustainability for forest health monitoring and triage: Case study of a mountain pine beetle (Dendroctonusponderosae)-impacted landscape
- Jose Negron, Joel McMillin, Carolyn H. Sieg, James F. Fowler, Kurt K. Allen, Linda L. Wadleigh, John A. Anhold, Ken E. Gibson. 2016. Variables associated with the occurrence of Ips beetles, red turpentine beetle and wood borers in live and dead ponderosa pines with post-fire injury
- Haifeng Zhou, Junyong Zhu, Roland L. Gleisner, Xueqing Qiu, Eric Horn, Jose Negron. 2016. Pilot-scale demonstration of SPORL for bioconversion of lodgepole pine to bioethanol and lignosulfonate
- M. M. Dudley, Jose Negron, N. A. Tisserat, W. D. Shepperd, W. R. Jacobi. 2015. Influence of climate on the growth of quaking aspen (Populus tremuloides) in Colorado and southern Wyoming
- Jose Negron, Ann M. Lynch, Willis C. Schaupp, Vladimir Bocharnikov. 2014. Douglas-fir tussock moth- and Douglas-fir beetle-caused mortality in a ponderosa pine/Douglas-fir forest in the Colorado Front Range, USA
- Daniel R. West, Jennifer S. Briggs, William R. Jacobi, Jose Negron. 2014. Mountain pine beetle-caused mortality over eight years in two pine hosts in mixed-conifer stands of the southern Rocky Mountains
- John M. Frank, William J. Massman, Brent E. Ewers, Laurie Huckaby, Jose Negron. 2014. Ecosystem CO2/H2O fluxes are explained by hydraulically limited gas exchange during tree mortality from spruce bark beetles
- Javier E. Mercado, Jose Negron. 2014. Revision of the new world species of Hylurgops LeConte, 1876 with the description of a new genus in the Hylastini (Coleoptera: Scolytinae) and comments on some Palearctic species
- Christopher J. Fettig, Kenneth E. Gibson, A. Steven Munson, Jose Negron. 2014. A comment on “Management for mountain pine beetle outbreak suppression: Does relevant science support current policy?"
- Jennifer G. Klutsch, Russell D. Beam, William R. Jacobi, Jose Negron. 2014. Bark beetles and dwarf mistletoe interact to alter downed woody material, canopy structure, and stand characteristics in northern Colorado ponderosa pine
- Ingrid Hoeger, Rolland Gleisner, Jose Negron, Orlando J. Rojas, Junyong Zhu. 2014. Mountain pine beetle-killed lodgepole pine for the production of submicron lignocellulose fibrils
- Javier E. Mercado, Richard W. Hofstetter, Danielle M. Reboletti, Jose Negron. 2014. Phoretic symbionts of the mountain pine beetle (Dendroctonus ponderosae Hopkins)
- Jose Negron, Christopher J. Fettig. 2014. Mountain pine beetle, a major disturbance agent in US western coniferous forests: A synthesis of the state of knowledge
- Nancy E. Gillette, David L. Wood, Sarah J. Hines, Justin B. Runyon, Jose Negron. 2014. The once and future forest: Consequences of mountain pine beetle treatment decisions
- Christopher J. Fettig, Kenneth E. Gibson, A. Steven Munson, Jose Negron. 2014. Cultural practices for prevention and control of mountain pine beetle infestations
- Sheryl L. Costello, William R. Jacobi, Jose Negron. 2013. Emergence of Buprestidae, Cerambycidae, and Scolytinae (Coleoptera) from mountain pine beetle-killed and fire-killed ponderosa pines in the Black Hills, South Dakota, USA
- Benjamin C. Bright, Andrew T. Hudak, Robert McGaughey, Hans-Erik Andersen, Jose Negron. 2013. Predicting live and dead tree basal area of bark beetle affected forests from discrete-return lidar
- Haifeng Zhou, Junyong Zhu, Xiaolin Luo, Shao-Yuan Leu, Xiaolei Wu, Roland L. Gleisner, Bruce S. Dien, Ronald E. Hector, Dongjie Yang, Xueqing Qiu, Eric Horn, Jose Negron. 2013. Bioconversion of Beetle-Killed Lodgepole Pine Using SPORL: Process Scale-up Design, Lignin Coproduct, and High Solids Fermentation without detoxification
- W. R. Jacobi, R. D. Koski, Jose Negron. 2013. Dutch elm disease pathogen transmission by the banded elm bark beetle Scolytus schevyrewi
- John R. Withrow, John Lundquist, Jose Negron. 2013. Spatial dispersal of Douglas-fir beetle populations in Colorado and Wyoming
- David J. P. Moore, Nicole A. Trahan, Phil Wilkes, Tristan Quaife, Britton B. Stephens, Kelly J. Elder, Ankur R. Desai, Jose Negron, Russell K. Monson. 2013. Persistent reduced ecosystem respiration after insect disturbance in high elevation forests
- Robert M. Hubbard, Charles C. Rhoades, Kelly J. Elder, Jose Negron. 2013. Changes in transpiration and foliage growth in lodgepole pine trees following mountain pine beetle attack and mechanical girdling
- Jose Negron. 2012. Historic forests and endemic mountain pine beetle and dwarf mistletoe
- Andrew T. Hudak, Ben Bright, Jose Negron, Robert McGaughey, Hans Andersen, Jeffrey A. Hicke. 2012. Predicting live and dead basal area in bark beetle-affected forests from discrete-return LiDAR
- Tania Schoennagel, Thomas T. Veblen, Jose Negron, Jeremy M. Smith. 2012. Effects of mountain pine beetle on fuels and expected fire behavior in lodgepole pine forests, Colorado, USA
- W. Matt Jolly, Russell A. Parsons, Ann M. Hadlow, Greg M. Cohn, Sara S. McAllister, John B. Popp, Robert M. Hubbard, Jose Negron. 2012. Relationships between moisture, chemistry, and ignition of Pinus contorta needles during the early stages of mountain pine beetle attack
- Sheryl L. Costello, Jose Negron, William R. Jacobi. 2011. Wood-boring insect abundance in fire-injured ponderosa pine
- Jose Negron. 2011. III. Insects
- Jose Negron, Sally J. McElwey, Jana C. Lee, Livy Williams, Jeffrey J. Witcosky, John B. Popp, Steve J. Seybold. 2011. Biology of the invasive banded elm bark beetle (Coleoptera: Scolytidae) in the western United States
- Junyong Zhu, Xiaolin Luo, Shen Tian, Roland L. Gleisner, Jose Negron, Eric Horn. 2011. Efficient ethanol production from beetle-killed lodgepole pine using SPORL technology and Saccharomyces cerevisiae without detoxification
- Jose Negron, Willis C. Schaupp, Lee Pederson. 2011. Flight periodicity of the Douglas-fir beetle, Dendroctonus pseudotsugae Hopkins (Coleoptera: Curculionidae: Scolytinae) in Colorado, U.S.A
- Jennifer G. Klutsch, Michael A. Battaglia, Daniel R. West, Sheryl L. Costello, Jose Negron. 2011. Evaluating potential fire behavior in lodgepole pine-dominated forests after a mountain pine beetle epidemic in north-central Colorado
- Carolyn H. Sieg, Joel McMillin, Kurt K. Allen, James F. Fowler, Jose Negron, Linda L. Wadleigh, John A. Anhold, Ken E. Gibson. 2010. Development of post-fire crown damage mortality thresholds in ponderosa pine
- Joel M. Egan, William R. Jacobi, Jose Negron, Sheri L. Smith, Daniel R. Cluck. 2010. Forest thinning and subsequent bark beetle-caused mortality in Northeastern California
- X. Luo, Roland L. Gleisner, S. Tian, Jose Negron, W. Zhu, E. Horn, X. J. Pan, Junyong Zhu. 2010. Evaluation of mountain beetle-infested lodgepole pine for cellulosic ethanol production by sulfite pretreatment to overcome recalcitrance of lignocellulose
- Barbara J. Bentz, Jacques Regniere, Christopher J. Fettig, Matt Hansen, Jane L. Hayes, Jeffrey A. Hicke, Rick G. Kelsey, Jose Negron, Steve J. Seybold. 2010. Climate change and bark beetles of the western United States and Canada: Direct and indirect effects
- Jana C. Lee, Shakeeb Hamud, Jose Negron, Jeffrey J. Witcosky, Steve J. Seybold. 2010. Semiochemical-mediated flight strategies of two invasive elm bark beetles: A potential factor in competitive displacement
- Matt Hansen, Jose Negron, A. Steven Munson, John A. Anhold. 2010. A retrospective assessment of partial cutting to reduce spruce beetle-caused mortality in the southern Rocky Mountains
- Jose Negron, Ken Gibson. 2009. Fire and bark beetle interactions
- Jose Negron, John B. Popp. 2009. The flight periodicity, attack patterns, and life history of Dryocoetes confusus Swaine (Coleoptera: Curculionida: Scolytinae), the western balsam bark beetle, in north central Colorado
- Paulius Zolubas, Jose Negron, A. Steven Munson. 2009. Modelling spruce bark beetle infestation probability
- Jennifer G. Klutsch, Jose Negron, Sheryl L. Costello, Charles C. Rhoades, Daniel R. West, John B. Popp, Rick Caissie. 2009. Stand characteristics and downed woody debris accumulations associated with a mountain pine beetle (Dendroctonus ponderosae Hopkins) outbreak in Colorado
- Barbara J. Bentz, Jesse Logan, Jim MacMahon, Craig D. Allen, Matt Ayres, Ed Berg, Allan Carroll, Matt Hansen, Jeff Hicke, Linda A. Joyce, Wallace Macfarlane, Steve Munson, Jose Negron, Tim Paine, Jim Powell, Ken Raffa, Jacques Regniere, Mary Reid, Bill Romme, Steve J. Seybold, Diana Six, Diana Tomback, Jim C. Vandygriff, Tom Veblen, Mike White, Jeff Witcosky, David Wood. 2009. Bark beetle outbreaks in western North America: Causes and consequences
- Janna C. Lee, Ingrid Aguayo, Ray Aslin, Gail Durham, Shakeeb Hamud, Beruce D. Moltzan, A. Steve Munson, Jose Negron, Travis Peterson, Iral R. Ragenovich, Jeffrey J. Witcosky, Steve J. Seybold. 2009. Co-occurrence of the invasive banded and European elm bark beetles (Coleoptera: Scolytidae) in North America
- Jose Negron, Joel D. McMillin, John A. Anhold, Dave Coulson. 2009. Bark beetle-caused mortality in a drought-affected ponderosa pine landscape in Arizona, USA
- Merrill R. Kaufmann, Gregory H. Aplet, Michael G. Babler, William L. Baker, Barbara J. Bentz, Michael G. Harrington, Brad C. Hawkes, Laurie Huckaby, Michael J. Jenkins, Daniel M. Kashian, Robert E. Keane, Dominik Kulakowski, Ward McCaughey, Charles W. McHugh, Jose Negron, John B. Popp, William H. Romme, Wayne Shepperd, Frederick W. Smith, Elaine K. Sutherland, Daniel Tinker, Thomas T. Veblen. 2008. The status of our scientific understanding of lodgepole pine and mountain pine beetles - a focus on forest ecology and fire behavior
- Jennifer G. Klutsch, Russell D. Beam, William R. Jacobi, Jose Negron. 2008. Fuel and stand characteristics in p. pine infested with mountain pine beetle, Ips beetle, and southwestern dwarf mistletoe in Colorado's Northern Front Range
- Joel McMillin, John Anhold, Jose Negron. 2008. Stand level impacts of Ips and Dendroctonus bark beetles in pine forest types of northern Arizona
- D. Allen-Reid, J. Anhold, D. Cluck, T. Eager, R. Mask, J. McMillin, S. Munson, Jose Negron, T. Rogers, D. Ryerson, E. Smith, S. Smith, B. Steed, R. Thier. 2008. Pinon pine mortality event in the Southwest: An update for 2005
- Jose Negron, Jill L. Wilson. 2008. Attributes associated with probability of infestation by the pinon Ips, Ips confusus, (Coleoptera: Scolytidae) in pinon pine, Pinus edulis
- Jose Negron, Barbara J. Bentz, Christopher J. Fettig, Nancy E. Gillette, Matt Hansen, Jane L. Hayes, Rick G. Kelsey, John Lundquist, Ann M. Lynch, Robert Progar, Steve J. Seybold. 2008. US Forest Service bark beetle research in the western United States: Looking toward the future
- Sheryl L. Costello, Jose Negron, William R. Jacobi. 2008. Traps and attractants for wood-boring insects in ponderosa pine stands in the Black Hills, South Dakota
- Jose Negron, Kurt Allen, Blaine Cook, John R. Withrow. 2008. Susceptibility of ponderosa pine, Pinus ponderosa (Dougl. Ex Laws.), to mountain pine beetle, Dendroctonus ponderosae Hopkins, attack in uneven-aged stands in the Black Hills of South Dakota and Wyoming USA
- Christopher J. Fettig, Kier D. Klepzig, Ronald f. Billings, A. Steven Munson, T. Evan Nebeker, Jose Negron, John Nowak. 2007. The effectiveness of vegetation management practices for prevention and control of bark beetle infestations in coniferous forests of the western and southern United States
- Jacek Hilszczanski, Wojciech Janiszewski, Jose Negron, A. Steve Munson. 2006. Stand characteristics and Ips typographus (L.) (Col., Curculionidae, Scolytinae) infestation during outbreak in northeastern Poland
- Carolyn H. Sieg, Joel D. McMillin, James F. Fowler, Kurt K. Allen, Jose Negron, Linda L. Wadleigh, John A. Anhold, Ken E. Gibson. 2006. Best predictors for postfire mortality of ponderosa pine trees in the Intermountain West
- Jose Negron, Kurt Allen, McMillin. Joel, Henry Burkwhat. 2006. Testing Verbenone for reducing mountain pine beetle attacks in ponderosa pine in the Black Hills, South Dakota
- Jose Negron, Jeffrey J. Witcosky, Robert J. Cain, James R. LaBonte, Donald A. Duerr, Sally J. McElwey, Jana C. Lee, Steve J. Seybold. 2005. The banded elm bark beetle: A new threat to elms in North America
- Jose Negron, Jill L. Wilson. 2003. Attributes associated with probability of infestation by the pinon ips, Ips confusus, (Coleoptera: Scolytidae) in pinon pine, Pinus edulis
- Anna W. Schoettle, Jose Negron. 2001. First report of two cone and seed insects on Pinus flexilis
- Jose Negron, John A. Anhold, A. Steve Munson. 2001. Within-stand spatial distribution of tree mortality caused by the Douglas-Fir beetle (Coleoptera: Scolytidae)
- Jose Negron, Wayne A. Shepperd, Steve A. Mata, John B. Popp, Lance A. Asherin, Anna W. Schoettle, John M. Schmid, David A. Leatherman. 2001. Solar treatments for reducing survival of mountain pine beetle in infested ponderosa and lodgepole pine logs
- John Lundquist, Jose Negron. 2000. Endemic forest disturbances and stand structure of ponderosa pine (Pinus ponderosa) in the Upper Pine Creek Research Natural Area, South Dakota, USA
- Jose Negron, Jill L. Wilson, John A. Anhold. 2000. Stand conditions associated with roundheaded pine beetle (Coleoptera: Scolytidae) infestations in Arizona and Utah
- Jose Negron, Willis C. Schaupp, Erik Johnson. 2000. Development and validation of a fixed-precision sequential sampling plan for estimating brood adult density of Dendroctonus pseudotsugae (Coleoptera: Scolytidae)
- Jose Negron, Willis C. Schaupp, Kenneth E. Gibson, John Anhold, Dawn Hansen, Ralph Thier, Phil Mocettini. 1999. Estimating extent of mortality associated with the Douglas-fir beetle in the Central and Northern Rockies
- Jose Negron. 1998. Probability of infestation and extent of mortality associated with the Douglas-fir beetle in the Colorado Front Range
- Jose Negron. 1997. Estimating probabilities of infestation and extent of damage by the roundheaded pine beetle in ponderosa pine in the Sacramento Mountains, New Mexico
- Brian W. Geils, John E. Lundquist, Jose Negron, Jerome S. Beatty. 1995. Disturbance regimes and their relationships to forest health.
- University of Puerto Rico, B.S., Biology, 1982
- Louisiana State University, Baton Rouge, M.S., Entomology, 1985
- Louisiana State University, Baton Rouge, Ph.D., Entomology, 1988
Last updated March 10, 2026