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Bret C. Harvey, PhD

Volunteer

Research Fish Biologist, Team Leader

Pacific Southwest Research Station

bret.harvey@usda.gov

Email

Publications & Products
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Education

Treesearch Publications

Bret C. Harvey, Jason L. White, Steven F. Railsback, Brian Staab, Daniel J. Isaak. 2026. Assessing the benefits of valley-bottom restoration for salmonids using spatially explicit, individual-based modeling
Steven F. Railsback, Bret C. Harvey. 2025. Including predation risk in mechanistic habitat assessment models for stream fish
Steven F. Railsback, Cara A. Gallagher, Volker Grimm, Matthew A. McCary, Bret C. Harvey. 2025. Empirical ecology to support mechanistic modelling: Different objectives, better approaches and unique benefits
David A. Roon, Jason B. Dunham, Joseph R. Benjamin, Bret C. Harvey, J. Ryan. Bellmore. 2025. Effects of riparian forest thinning on resident salmonid fishes in coastal northern California catchments
Steven F. Railsback, Bret C. Harvey. 2025. Representing mortality risk in mechanistic models
Daniel Ayllón, Steven F. Railsback, Bret C. Harvey, Graciela G. Nicola, Benigno Elvira, Ana Almodóvar. 2025. Behavioural plasticity in circadian foraging patterns increases resistance of brown trout populations to environmental change
Andrew P. Kinziger, Steven R. Fong, Rodney J. Nakamoto, Jason L. White, Bret C. Harvey. 2024. Contemporary evolution of neutral genetic structure in introduced Klamath smallscale suckers (Catostomus rimiculus) recapitulates native patterns
Bret C. Harvey, Jason L. White, Rodney J. Nakamoto, Steven F. Railsback. 2024. Empirical and model‐based evaluation of a step‐pool stream restoration project: Consequences for a highly valued fish population
Steven F. Railsback, Bret C. Harvey. 2023. Can thermal refuges save salmonids? Simulation of cold‐pool benefits to salmonid populations
Steven F. Railsback, Bret C. Harvey, Daniel Ayllón. 2023. InSTREAM 7 user manual: model description, software guide, and application guide
David A. Roon, Jason B. Dunham, J. Ryan Bellmore, Deanna H. Olson, Bret C. Harvey. 2022. Influence of riparian thinning on trophic pathways supporting stream food webs in forested watersheds
Bret C. Harvey, Rodney J. Nakamoto, Adam J. R. Kent, Christian E. Zimmerman. 2021. The distribution of anadromy and residency in steelhead/rainbow trout in the Eel River, northwestern California
Steven F. Railsback, Daniel Ayllón, Bret C. Harvey. 2021. InSTREAM 7: Instream flow assessment and management model for stream trout
Andrew P. Kinziger, Jason L. White, Rodney J. Nakamoto, Bret C. Harvey. 2021. Recent, small beginnings: genetic analysis suggests Catostomus rimiculus (Klamath smallscale sucker) in the Smith River, California, are introduced
Steven F. Railsback, Bret C. Harvey, Daniel Ayllón. 2021. Importance of the daily light cycle in population–habitat relations: A simulation study
Bret C. Harvey, Steven F. Railsback. 2021. All fish, all the time: A good general objective for fish passage projects?
Steven F. Railsback, Uta Berger, Jarl Giske, George I Hagstrom, Bret C. Harvey, Christina Semeniuk, Volker Grimm. 2020. Bridging levels from individuals to communities and ecosystems: including adaptive behavior and feedbacks in ecological theory and models
Steven Railsback, Bret C. Harvey, Daniel Ayllón. 2020. Contingent trade-off decisions with feedbacks in cyclical environments: testing alternative theories
Daniel Ayllón, Steven F. Railsback, Bret C. Harvey, Inmaculada García Quirós, Graciela G. Nicola, Benigno Elvira, Ana Almodóvar. 2019. Mechanistic simulations predict that thermal and hydrological effects of climate change on Mediterranean trout cannot be offset by adaptive behaviour, evolution, and increased food production
Andrew P. Kinziger, Rodney J. Nakamoto, Andy Aguilar, Bret C. Harvey. 2019. California roach (Hesperoleucus symmetricus) in the Eel River of northwestern California: native or introduced?
David Roon, Jason Dunham, Bret C. Harvey, J. Ryan Bellmore, Deanna H. Olson, Gordon Reeves. 2017. Evaluating the ecological trade-offs of riparian thinning for headwater stream ecosystems in second-growth redwood forests
Jason L. White, Bret C. Harvey. 2017. Response of steelhead/rainbow trout (Oncorhynchus mykiss) populations to debris flows
Valery E. Forbes, Chris J. Salice, Bjorn Birnir, Randy J.F. Bruins, Peter Calow, Virginie Ducrot, Nika Galic, Kristina Garber, Bret C. Harvey, Henriette Jager, Andrew Kanarek, Robert Pastorok, Steve F. Railsback, Richard Rebarber, Pernille Thorbek. 2017. A framework for predicting impacts on ecosystem services from (sub)organismal responses to chemicals
Bret C. Harvey, Jason L. White. 2017. Axes of fear for stream fish: water depth and distance to cover
Bret C. Harvey, Jason L. White. 2016. Use of cover for concealment behavior by Rainbow Trout: influences of cover structure and area
Steven F. Railsback, Bret C. Harvey, Sarah J. Kupferberg, Margaret M. Lang, Scott McBain, Hart Welsh. 2016. Modeling potential river management conflicts between frogs and salmonids
F. K. Ligon, Rodney J. Nakamoto, Bret C. Harvey, P. F. Baker. 2016. Use of streambed substrate as refuge by steelhead or rainbow trout Oncorhynchus mykiss during simulated freshets
Steven F. Railsback, Bret C. Harvey, Jason L. White. 2015. Effects of spatial extent on modeled relations between habitat and anadromous salmonid spawning success
Steven F. Railsback, Bret C. Harvey, Jason L. White. 2014. Facultative anadromy in salmonids: linking habitat, individual life history decisions, and population-level consequences
Andrew P. Kinziger, Rodney J. Nakamoto, Bret C. Harvey. 2014. Local-scale invasion pathways and small founder numbers in introduced Sacramento pikeminnow (Ptychocheilus grandis)
Bret C. Harvey, Jason L. White, Rodney J. Nakamoto, Steven F. Railsback. 2014. Effects of streamflow diversion on a fish population: combining empirical data and individual-based models in a site-specific evaluation
Bret C. Harvey, Steve Railsback. 2014. Feeding modes in stream salmonid population models: Is drift feeding the whole story?
Bret C. Harvey, Rodney J. Nakamoto. 2013. Seasonal and among-stream variation in predator encounter rates for fish prey
S. F. Railsback, M. Gard, Bret C. Harvey, Jason L. White, J.K.H. Zimmerman. 2013. Contrast of degraded and restored stream habitat using an individual-based salmon model
Steven F. Railsback, Bret C. Harvey. 2013. Trait-mediated trophic interactions: is foraging theory keeping up?
Jose Ricardo Barradas, Lucas G. Silva, Bret C. Harvey, Nelson F. Fontoura. 2012. Estimating migratory fish distribution from altitude and basin area: a case study in a large Neotropical river
Brian C. Spence, Walter G. Duffy, John Carlos Garza, Bret C. Harvey, Susan M. Sogard, Laurie A. Weitkamp, Thomas H. Williams, David A. Boughton. 2011. Historical occurrence of coho salmon (Oncorhynchus kisutch) in streams of the Santa Cruz Mountain region of California: response to an Endangered Species Act petition to delist coho salmon south of San Francisco Bay
Andrew Kinziger, Rodney J. Nakamoto, Eric Anderson, Bret C. Harvey. 2011. Small founding number and low genetic diversity in an introduced species exhibiting limited invasion success (speckled dace, Rhinichthys ocsulus)
Bret C. Harvey, Steven Railsback. 2011. Effects of passage barriers on demographics and stability properties of a virtual trout populations
Steven Railsback, Bret C. Harvey. 2011. Importance of fish behaviour in modelling conservation problems: food limitation as an example
Bret C. Harvey, Margaret A. Wilzbach. 2010. Carcass addition does not enhance juvenile salmonid biomass, growth, or retention in six Northwestern California streams.
Brian H. Hill, Frank H. McCormick, Bret C. Harvey, Sherri Johnson, Mel Warren, Colleen M. Elonen. 2010. Microbial enzyme activity, nutrient uptake and nutrient limitation in forested streams
Bret C. Harvey, Steven F. Railsback. 2009. Exploring the persistence of stream-dwelling trout populations under alternative real-world turbidity regimes with an individual-based model
Bret C. Harvey, Jason L. White, Rodney J. Nakamoto. 2009. The effect of deposited fine sediment on summer survival and growth of rainbow trout in riffles of a small stream
Steven F. Railsback, Bret C. Harvey, Stephen K. Jackson, Roland H. Lamberson. 2009. InSTREAM: the individual-based stream trout research and environmental assessment model
Bret C. Harvey, Jason L. White. 2008. Use of benthic prey by salmonids under turbid conditions in a laboratory stream
Jason L. White, Bret C. Harvey. 2007. Winter feeding success of stream trout under different streamflow and turbidity conditions
Bret C. Harvey, Steven F. Railsback. 2007. Estimating multi-factor cumulative watershed effects on fish populations with an individual-based model
Bret C. Harvey, Rodney J. Nakamoto, Jason L. White. 2006. Reduced streamflow lowers dry-season growth of rainbow trout in a small stream
Steven F. Railsback, Bret C. Harvey, John W. Hayse, Kirk E. Lagory. 2005. Tests of a theory for diel activity and habitat selection
Margaret A. Wilzbach, Bret C. Harvey, Jason L. White, Rodney J. Nakamoto. 2005. Effects of riparian canopy opening and salmon carcass addition on the abundance and growth of resident salmonids
Bret C. Harvey, Jason L. White, Rodney J. Nakamoto. 2005. Habitat-specific biomass, survival and growth of rainbow trout (Oncorhynchus mykiss) during summer in a small coastal stream
Bret C. Harvey, Jason L. White, Rodney J. Nakamoto. 2004. An emergent multiple predator effect may enhance biotic resistance in a stream fish assemblage
Jason L. White, Bret C. Harvey. 2003. Basin-scale patterns in the drift of embryonic and larval fishes and lamprey ammocoetes in two coastal rivers
Bret C. Harvey, Jason L. White, Rodney J. Nakamoto. 2002. Habitat relationships and larval drift of native and nonindigenous fishes in neighboring tributaries of a coastal California river
Carl D. Reese, Bret C. Harvey. 2002. Temperature-dependent interactions between juvenile steelhead and Sacramento pikeminnow in laboratory streams
Steven F. Railsback, Bret C. Harvey, Roland H. Lamberson, Derek E. Lee, Claasen Nathan J., Shuzo Yoshihara. 2002. Population-level analysis and validation of an individual-based cutthroat trout model
Steven F. Railsback, Bret C. Harvey. 2002. Analysis of habitat-selection rules using an individual-based model
Steven F. Railsback, Bret C. Harvey. 2001. Individual-based model formulation for cutthroat trout, Little Jones Creek, California
Jason L. White, Bret C. Harvey. 2001. Effects of an introduced piscivorous fish on native benthic fishes in a coastal river
Hartwell H. Welsh, Garth R. Hodgson, Bret C. Harvey, Maureen F. Roche. 2001. Distribution of juvenile coho salmon in relation to water temperatures in tributaries of the Mattole River, California
Bret C. Harvey, Rodney J. Nakamoto, Jason L. White. 1999. The influence of large woody debris and a bankfull flood on movement of adult resident coastal cutthroat trout (Oncorhynchus clarki) during fall and winter
Steven F. Railsback, Roland H. Lamberson, Bret C. Harvey, Walter E. Duffy. 1999. Movement rules for individual-based models of stream fish
Bret C. Harvey, Thomas E. Lisle. 1999. Scour of chinook salmon redds on suction dredge tailings
Bret C. Harvey, Rodney J. Nakamoto. 1999. Diel and seasonal movements by adult Sacramento pikeminnow (Ptychocheilus grandis) in the Eel River, northwestern California
Jason L. White, Bret C. Harvey. 1999. "Habitat separation of prickly sculpin, Cottus asper, and coastrange sculpin, Cottus aleuticus, in the mainstem Smith River, northwestern California"
Jason L. White, Bret C. Harvey. 1999. Habitat separation of prickly sculpin, Cottus asper, and coastrange sculpin, Cottus aleuticus, in the mainstem Smith River, northwestern California
Bret C. Harvey, Thomas E. Lisle. 1998. Effects of suction dredging on streams: a review and an evaluation strategy
Bret C. Harvey. 1998. The influence of large woody debris on retention, immigration, and growth of coastal cutthroat trout (Oncorhynchus clarki clarki) in stream pools
Bret C. Harvey, Rodney J. Nakamoto. 1997. Habitat-dependent interactions between two size-classes of juvenile steelhead in a small stream
Bret C. Harvey, Rodney J. Nakamoto. 1996. Effects of steelhead density on growth of Coho salmon in a small coastal California stream

University of Oklahoma, Ph.D., Zoology, 1987
University of California, M.S., Ecology, 1982
University of California, B.S., Wildlife and Fisheries Biology, 1980

Last updated March 10, 2026

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