James Hanula, PhD

Treesearch Publications

• Michael Ulyshen, Scott Horn, James Hanula. 2022. Decadal patterns of forest and pollinator recovery following the eradication of an invasive shrub.
• Michael Ulyshen, Scott Horn, James Hanula. 2020. Effects of Chinese privet on bees and their vertical distribution in riparian forests
• Stephen Fraedrich, Thomas Harrington, Qiong Huang, Stan Zarnoch, James Hanula, Susan Best. 2018. Brood production by xyleborus glabratus in bolts from trees infected and uninfect ed with the laurel wilt pathogen, raffaelea lauricola.
• J. Kalina, S.K. Braman, James Hanula. 2017. Host utilization of Chinese Privet (Lamiales: Oleaceae) and host choice by Leptoypha mutica (Hemiptera: Tingidae)
• J. Kalina, S.K. Braman, James Hanula. 2017. Developmental biology of Leptoypha mutica (Hemiptera: Tingidae) on Chinese Privet (Lamiales: Oleaceae)
• Yanzhuo Zhang, James Hanula, Scott Horn, Cera Jones, S. Kristine Braman, Jianghua Sun. 2016. Fundamental host range of Leptoypha hospita (Hemiptera: Tingidae), a potential biological control agent of Chinese privet
• James Hanula, Michael Ulyshen, Scott Horn. 2016. Conserving pollinators in North American forests: A review
• James Hanula, Albert (Bud) E. Mayfield, Laurie S. Reid, Scott Horn. 2016. Influence of trap distance from a source population and multiple traps on captures and attack densities of the redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae)
• R. Scott Cameron, James Hanula, Stephen Fraedrich, Chip Bates. 2015. Progression and Impact of Laurel Wilt Disease within Redbay and Sassafras Populations in Southeast Georgia
• M. A. Hughes, J.A. Smith, R. C. Ploetz, P. E. Kendra, Albert (Bud) E. Mayfield, James Hanula, J. Hulcr, L.L. Stelinski, S. Cameron,  J. J. Riggins, D. Carrillo, R. Rabaglia, J. Eickwort. 2015. Recovery plan for laurel wilt on redbay and other forest species caused by Raffaelea lauricola and disseminated by Xyleborus glabratus
• James Hanula, Scott Horn, Joseph J. O'Brien. 2015. Have changing forests conditions contributed to pollinator decline in the southeastern United States?
• James Hanula, Albert (Bud) E. Mayfield. 2014. Redbay Ambrosia Beetle (xyleborus glabratus eichoff) (coleoptera: curculionidae)
• C.E. Jones, Nathan P. P. Havill, James Hanula, S.K. Braman. 2014. Post release recovery of hemlock woolly adelgid predators in the North Georgia mountains.
• C.E. Jones, James Hanula, S.K. Braman. 2014. Emergence of Laricobius nigrinus (Fender) (Coleoptera: Derodontidae) in the North Georgia mountains.
• Joshua W. Lobe, Mac A. Callaham Jr., Paul F. Hendrix, James Hanula. 2014. Removal of an invasive shrub (Chinese privet: Ligustrum sinense Lour) reduces exotic earthworm abundance and promotes recovery of native North American earthworms
• Albert (Bud) E. Mayfield, Barbara C. Reynolds, Carla I. Coots, Nathan P. P. Havill, Cavell Brownie, Andrew R. Tait, James Hanula, Shimat V. Joseph, Ashley B. Galloway. 2014. Establishment, hybridization and impact of Laricobius predators on insecticide-treated hemlocks: Exploring integrated management of the hemlock woolly adelgid
• M. Lake Maner, James Hanula, Scott Horn. 2014. Population trends of the redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae): does utilization of small diameter redbay trees allow populations to persist?
• C.W. Johnson, R.S. Cameron, James Hanula, C. Bates. 2014. The attractiveness of manuka oil and ethanol, alone and in combination, to xyleborus glabratus (Coleoptera: Curculionidae: Scolytinae) and other curculionidae
• Jacob R. Hudson, James Hanula, Scott Horn. 2014. Impacts of removing Chinese privet from riparian forests on plant communities and tree growth five years later
• Jacob R. Hudson, James Hanula, Scott Horn. 2014. Removing Chinese privet from riparian forests still benefits pollinators five years later
• M. Lake Maner, James Hanula, S. Kristine Braman. 2014. Rearing redbay ambrosia beetle, Xyleborus glabratus (Coleoptera: Curculionidae: Scolytinae), on semi-artifical media
• Yanzhuo Zhang, James Hanula, Joseph J. O'Brien, Scott Horn, Braman Kris, Jianghua Sun. 2013. Evaluation of the impacts of herbivory by lace bugs on Chinese privet (Ligustrum sinense) survival and physiology
• M. Lake Maner, James Hanula, S. Kristine Braman. 2013. Gallery productivity, emergence, and flight activity of the redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae)
• M. Lake Maner, James Hanula, S. Kristine Braman. 2013. Evaluation of screen barriers on redbay trees to protect them from Xyleborus glabratus (Coleoptera: Curculionidae: Scolytinae) and distribution of initial attacks in relation to stem moisture content, diameter, and height
• James Hanula, Brian Sullivan, David Wakarchuk. 2013. Variation in manuka oil lure efficacy for capturing Xyleborus glabratus (Coleoptera: Curculionidae: Scholytinae), and Cubeb oil as an alternative attractant
• Michael Ulyshen, James Hanula, Robert L. Blinn, Gene Kritsky. 2012. Saproxylic Hemiptera Habitat Associations
• Michael Ulyshen, Scott Horn, James Hanula. 2012. Growth responses of mature loblolly pine to dead wood manipulations
• Jordon L. Burke, James Hanula, Scott Horn, Jackson P. Audley, Kamal JK. Gandhi. 2012. Efficacy of two insecticides for protecting loblolly pines (Pinus taeda L.) from subcortical beetles (Coleoptera: Curculionidae and Cerambycidae)
• James Hanula, Michael Ulyshen, Dale D. Wade. 2012. Impacts of prescribed fire frequency on coarse woody debris volume, decomposition and termite activity in the longleaf pine flatwoods of Florida
• Albert (Bud) E. Mayfield, James Hanula. 2012. Effect of tree species and end seal on attractiveness and utility of cut bolts to the redbay Ambrosia beetle and granulate Ambrosia beetle (Coleoptera: Curculionidae: Scolytinae)
• Christopher E. Moorman, Liessa T. Woen, John Kilgo, James Hanula, Scott Horn, Michael Ulyshen. 2012. Arthropod abundance and seasonal bird use of bottomland forest harvest gaps
• Yanzhuo Zhang, James Hanula, Scott Horn. 2012. The biology and preliminary host range of Megacopta cribraria (Heteroptera: Plataspidae) and its impact on kudzu growth
• Shimat V. Joseph, S. Kristine Braman, Jim Quick, James Hanula. 2011. The range and response of Neonicotinoids on hemlock woolly adelgid, Adelges tsugae (Hemiptera: Adelgidae)
• Michael Ulyshen, Villu Soon, James Hanula. 2011. Vertical distribution and seasonality of predatory wasps (Hymenoptera: Vespidae) in a temperate deciduous forest
• S.V. Joseph, James Hanula, S.K. Braman. 2011. Distribution and Abundance of Adelges tsugae (Hemiptera: Adelgidae) Within Hemlock Trees
• Yanzhuo Zhang, James Hanula, Scott Horn, Kristine Braman, Jianghua Sun. 2011. Biology of Leptoypha hospita (Hemiptera: Tingidae), a Potential Biological Control Agent of Chinese Privet
• James Hanula, Scott Horn. 2011. Removing an invasive shrub (Chinese privet) increases native bee diversity and abundance in riparian forests of the southeastern United States
• Michael Ulyshen, Thomas Pucci, James Hanula. 2011. The importance of forest type, tree species and wood posture to saproxylic wasp (Hymenoptera) communities in the southeastern United States.
• James Hanula, Scott Horn. 2011. Removing an exotic shrub from riparian forests increases butterfly abundance and diversity
• S. V. Joseph, James Hanula, S. K. Braman, F. J. Byrne. 2011. Effects of fertilizer and low rates of Imidacloprid on Adelges tsugae (Hemiptera: Adelgidae).
• James Hanula, Michael Ulyshen, Scott Horn. 2011. Effect of trap type, trap position, time of year, and beetle density on captures of the Redbay Ambrosia Beetle (Coleoptera: Curculionidae: Scolytinae).
• Scott Horn, James Hanula. 2011. Influence of trap color on collection of the recently introduced Bean Plataspid, Megacopta cribraria (Hemiptera: Plataspidae).
• Yan-Zhuo Zhang, James Hanula, Sun Jiang-Hua. 2011. Herbivory effects of Argopistes tsekooni, a chrysomelid beetle, on container-grown Chinese privet, Ligustrum sinense
• Melissa Churchel, James Hanula, C. Wayne Berisford, James Vose, Mark Dalusky. 2011. Impact of Imidacloprid for control of Hemlock Woolly Adelgid on nearby aquatic macroinvertebrate asseblages.
• S.V. Joseph, James Hanula. 2011. Effects of fertilization of four hemlock species on Adelges tsugae (Hemiptera: Adelgidae) growth and feeding preference of predators.
• Michael Ulyshen, James Hanula. 2010. Host-use patterns of Eriotremex formosanus (Hymenoptera: Siricidae) in South Carolina, U.S.A.
• Michael Ulyshen, Scott Horn, James Hanula. 2010. First Southeast U.S. records for Lacconotus punctatus Leconte (Coleoptera: Mycteridae), an early seasonal canopy specialist?
• Michael Ulyshen, Villa Soon, James Hanula. 2010. On the vertical distribution of bees in a temperate deciduous forest
• Michael D. Ulyshen, Scott Horn, James Hanula. 2010. Response of beetles (Coleoptera) at three heights to the experimental removal of an invasive shrub, Chinese privet (Lingustrum sinense), from floodplain forests
• Michael Ulyshen, James Hanula. 2010. Patterns of saproxylic beetle succession in loblolly pine.
• James Hanula, John Taylor. 2010. Controlling chinese privet can be a grind.
• James Hanula, Scott Horn, John W. Taylor. 2010. Chinese Privet (Ligustrum sinense) removal and its effect on native plant communities of Riparian Forests
• Michael D. Ulyshen, James Hanula. 2009. Responses of arthropods to large-scale manipulations of dead wood in loblolly pine stands of the Southeastern United States
• Michael Ulyshen, James Hanula. 2009. Responses of Arthropods to large scale manipulations of dead wood in loblolly pine stands of the southeastern United States
• James Hanula, Scott Horn. 2009. Impact of Chinese privet and its removal on pollinator diversity and abundance
• James Hanula, Dale Wade, Joseph O'Brien, Susan Loeb. 2009. Ground-dwelling arthropod association with coarse woody debris following long-term dormant season prescribed burning in the longleaf pine flatwoods of North Florida
• Michael Ulyshen, James Hanula. 2009. Habitat associations of saproxylic beetles in the southeastern United States: A comparison of forest types, tree species and wood postures.
• Michael D. Ulyshen, James Hanula. 2009. Litter-dwelling arthropod abundance peaks near coarse woody debris in loblolly pine forests of the southeastern United States
• M.D. Ulyshen, James Hanula. 2009. Habitat associations of saproxylic beetles in the southeastern United States: A comparison of forest types, tree species and wood postures.
• Michael D. Ulyshen, James Hanula. 2009. Litter-dwelling arthropod abundance peaks near coarse woody debris in loblolly pine forest of the southeastern United States.
• Y-Z Zhang, J. Sun, James Hanula. 2009. Biology and life history of Argopistes tsekooni (Coleoptera: Chrysomelidae) in China, a promising biological control agent of Chinese privet.
• James Hanula, D.D. Wade, Joseph J. O'Brien, Susan Loeb. 2009. Ground-dwelling arthropod association with coarse woody debris following long-term dormant season prescribed burning in the longleaf pine flatwoods of north Florida
• Yan Zhuo Zhang, James Hanula, Jiang Hua Sun. 2008. Host Specificity of Argopistes tsekooni (Coleoptera: Chrysomelidae), a Potential Biological Control Agent of Chinese Privet
• Y-Z Zhang, James Hanula, J. Sun. 2008. Survey for potential insect biological control agents of Ligustrum sinense (Scrophulariales: Oleaceae) in China.
• Joshua W. Campbell, James Hanula, Thomas A. Waldrop. 2008. Effects of prescribed fire and fire surrogates on Saproxylic coleoptera in the Southern Appalachians of North Carolina
• James Hanula, Brian Sullivan. 2008. Manuka oil and phoebe oil are attractive baits for xyleborus glabratus (coleoptera: scolytinae), the vector of Laurel Wilt
• James Hanula, Albert E. Mayfield, Stephen Fraedrich, Robert J. Babaglia. 2008. Biology and host associations of redbay ambrosia beetle (Coleoptera: Curculionidae: Scolytinae), exotic vector of laurel wilt killing redbay trees in the Southeastern United States
• Scott Horn, James Hanula. 2008. Relationship of coarse woody debris to arthropod availability for red-cockaded woodpeckers and other bark-foraging birds on loblolly pine boles
• Joshua W. Campbell, James Hanula, Kenneth Outcalt. 2008. Effects of prescribed fire and other plant community restoration treatments on tree mortality, bark beetles, and other saproxylic coleoptera of longleaf pine, Pinus palustris Mill., on the coastal plain of Alabama
• Michael Ulyshen, James Hanula. 2007. A comparison of the Beetle (Coleoptera) Fauna Captured at two heights above the ground in a North American temperate deciduous forest
• John Klepac, Robert Rummer, James Hanula, Scott Horn. 2007. Mechanical removal of Chinese privet
• John W. Campbell, James Hanula, Thomas A. Waldrop. 2007. Observations of Speyeria diana (Diana Fritillary) utilizing forested areas in North Carolina that have been Mechanically thinned and burned
• Michael D. Ulyshen, James Hanula, Scott Horn. 2007. Burying beetles (Coleoptera: Silphidae) in the forest canopy: The unusual case of Nicrophorus pustulatus Herschel
• Christopher E. Moorman, Liessa T. Bowen, John C. Kilgo, Clyde E. Sorenson, James Hanula, Scott Horn, Mike D. Ulyshen. 2007. Seasonal diets of insectivorous birds using canopy gaps in a bottomland forest
• Michael D. Ulyshen, James Hanula. 2007. A comparison of the beetle (Coleoptera) fauna captured at two heights above the ground in a North American temperate deciduous forest
• Scott Horn, James Hanula. 2006. Burlap bands as a sampling technique for green anoles (Anolis carolinensis) and other reptiles commonly found in tree boles
• Simon J. Grove, James Hanula. 2006. Insect biodiversity and dead wood: proceedings of a symposium for the 22^nd International Congress of Entomology
• Scott Horn, James Hanula. 2006. Burlap bands as a sampling technique for green anoles (Anolis carolinensis) and other reptiles commonly found on tree boles
• James Hanula, Scott Horn, Dale D. Wade. 2006. The role of dead wood in maintaining arthropod diversity on the forest floor
• Michael D. Ulyshen, James Hanula, Scott Horn, John C. Kilgo, Christopher E. Moorman. 2005. The response of ground beetles (Coleoptera: Carabidae) to selection cutting in a South Carolina bottomland hardwood forest
• Michael D. Ulyshen, James Hanula, Scott Horn. 2005. Using Malaise traps to sample ground beetles (Coleoptera: Carabidae)
• Scott Horn, James Hanula, Michael D. Ulyshen, John C. Kilgo. 2005. Abundance of green tree frogs and insects in artificial canopy gaps in a bottomland hardwood forest
• Michael D. Ulyshen, James Hanula, Scott Horn, John C. Kilgo, Christopher E. Moorman. 2005. Herbivorous insect response to group selection cutting in a southeastern bottomland hardwood forest
• Kurtis R. Moseley, Steven B. Castleberry, James Hanula, W. Mark Ford. 2004. Diet of southern toads (Bufo terrestris) in loblolly pine (Pinus taeda) stands subject to coarse woody debris manipulations
• Michael D. Ulyshen, James Hanula, Scott Horn, John C. Kilgo, Christopher E. Moorman. 2004. Spatial and temporal patterns of beetles associated with coarse woody debris in managed bottomland hardwood forests
• Alex C. Mangini, William W. Bruce, James Hanula. 2004. Radiographic Analysis of Shortleaf Pine Seeds From the Ouachita and Ozark National Forests
• Scott Horn, James Hanula. 2004. Impact of Seed Predators on the Herb Baptisia Lanceolata (Fabales: Fabaceae)
• James Hanula, Scott Horn. 2004. Availability and abundance of prey for the red-cockaded woodpecker
• Michael D. Ulyshen, James Hanula. 2004. Diversity and Seasonal Activity of Carrion Beetles (Coleoptera: Silphidae) in Northeastern Georgia
• Scott Horn, James Hanula. 2004. A Survey of Cavity-Nesting Bees and Wasps in Loblolly Pine Stands of the Savannah River Site, Aiken County, South Carolina
• Christopher Asaro, Susan Loeb, James Hanula. 2003. Cone consumption by southeastern fox squirrels: A potential basis for clonal preferences in a loblolly and slash pine seed orchard
• Nenad Mihelcic, James Hanula, Gary L. DeBarr. 2003. Pathogenic Microorganisms Associated With the Southern Pine Coneworm (Lepidoptera: Pyralidae) Attacking Loblolly Pine
• James Hanula, Dale D. Wade. 2003. Influence of long-term dormant-season burning and fire exclusion on ground-dwelling arthropod populations in longleaf pine flatwoods ecosystems
• Scott Horn, James Hanula. 2002. Comparison of arthropod prey of red-cockaded woodpeckers on the boles of long-leaf and loblolly pines
• Scott Horn, James Hanula. 2002. Life History and Habitat Associations of the Broad Wood Cockroach, Purcoblatta lata (Blattaria: Blattellidae) and Other Native Cockroaches in the Coastal Plain of South Carolina
• Ellen A. Gray, John C. Rennie, Thomas A. Waldrop, James Hanula. 2002. Patterns of Seed Productions in Table Mountain Pine
• James Hanula, James R. Meeker, Daniel R. Miller, Edward L. Barnard. 2002. Association of wildfire with tree health and numbers of pine bark beetles, reproduction weevils and their associates in Florida
• Theresa Pitts-Singer, James Hanula, Joan L. Walker. 2002. Insect Pollinators of Three Rare Plants in a Florida Longleaf Pine Forest
• James Hanula, Gary L. DeBarr, Julie C. Weatherby, Larry R. Barber, C. Wayne Berisford. 2002. Degree-day model for timing insecticide applications to control Dioryctria amatella (Lepidoptera: Pyralidae) in loblolly pine seed orchards
• James Hanula, Dale D. Wade. 2000. Effects of 40 years of winter burning in longleaf pine on insects and other arthropods
• James Hanula, Kathleen Franzreb, William D Pepper. 2000. Longleaf pine characteristics associated with arthropods available for red-cockaded woodpeckers
• James Hanula, Donald Lipscomb, Kathleen Franzreb, Susan Loeb. 2000. Diet of nestling red-cockaded woodpeckers at three locations
• Gary L. DeBarr, James Hanula, Christine G. Niwa, John C Nord. 2000. Synthetic pheromones disrupt male Dioryctria spp. moths in a loblolly pine seed orchard
• James Hanula, R. Todd Engstrom. 2000. Comparison of red-cockaded woodpecker (Picoides borealis) nestling diet in old-growth and old-field longleaf pine (Pinus palustris) habitats
• Kirsten C. New, James Hanula. 1998. Effect of time elapsed after prescribed burning in longleaf pine stands on potential prey of the red-cockaded woodpecker
• James Hanula, Kirsten C.P. New. 1996. A Trap For Capturing Arthropods Crawling up Tree Boles
• James Hanula, Kathleen Franzreb. 1995. Arthropod prey of nestling red-cockaded woodpeckers in the upper coastal plain of South Carolina

• University of Georgia, Ph.D., Forest Entomology, 1983
• University of Georgia, M.S., Forest Entomology, 1981
• Texas A&M University, B.S., Forest Management, 1978