John J. Keane, PhD

Treesearch Publications

• Kristin M. Brunk, Joshua F. Goldberg, Charles Maxwell, M. Zachariah Peery, Gavin M. Jones, Lief V. Gallagher, H. Anu Kramer, Anthony LeRoy Westerling, John J. Keane, Stefan Kahl, Connor M. Wood. 2025. Bioregional-scale acoustic monitoring can support fire-prone forest restoration planning
• Kate McGinn, Ceeanna Zulla, Marilyn Wright, Zachary Wilkinson, Brian Dotters, Kevin Roberts, John J. Keane, M. Zachariah Peery, Gavin M. Jones. 2024. Energetics explain predator occurrence and movement in pyrodiverse landscapes
• Whitney A. Watson, Daniel F. Hofstadter, Gavin M. Jones, H. Anu Kramer, Nicholas F. Kryshak, Ceeanna J. Zulla, Sheila A. Whitmore, Virginia O’Rourke, John J. Keane, R. J. Gutierrez, M. Zachariah Peery. 2024. Characterizing juvenile dispersal dynamics of invasive Barred Owls: Implications for management
• Whitney Watson, Connor Wood, Kevin Kelly, Daniel Hofstadter, Nicholas Kryshak, Ceeanna Zulla, Sheila Whitmore, Virginia O’Rourke, John J. Keane, M Zachariah. Peery. 2023. Passive acoustic monitoring indicates Barred Owls are established in northern coastal California and management intervention is warranted
• Corbin C. Kuntze, Jonathan N. Pauli, Ceeanna J. Zulla, John J. Keane, Kevin N. Roberts, Brian P. Dotters, Sarah Sawyer, M. Zachariah. Peery. 2023. Landscape heterogeneity provides co‐benefits to predator and prey
• Zachary A. Wilkinson, H. Anu Kramer, Gavin M. Jones, Ceeanna J. Zulla, Kate McGinn, Josh M. Barry, Sarah Sawyer, Richard Tanner, R. J. Gutierrez, John J. Keane, Zachariah Peery. 2023. Tall, heterogeneous forests improve prey capture, delivery to nestlings, and reproductive success for Spotted Owls in southern California
• Ceeanna J. Zulla, Gavin M. Jones, H. Anu Kramer, John J. Keane, Kevin N. Roberts, Brian P. Dotters, Sarah Sawyer, Sheila A. Whitmore, William J. Berigan, Kevin G. Kelly, R. J. Gutierrez, M. Zachariah Peery. 2023. Forest heterogeneity outweighs movement costs by enhancing hunting success and reproductive output in California spotted owls
• Kate A. McGinn, Benjamin Zuckerberg, Jonathan N. Pauli, Ceeanna J. Zulla, William J. Berigan, Zachary A. Wilkinson, Josh M. Barry, John J. Keane, R. J. Gutiérrez, M. Zachariah. Peery. 2023. Older forests function as energetic and demographic refugia for a climate-sensitive species
• Kevin G. Kelly, Connor M. Wood, Kate McGinn, H. Anu Kramer, Sarah Sawyer, Sheila Whitmore, Dana Reid, Stefan Kahl, Aimee Reiss, Jonathan Eiseman, William Berigan, John J. Keane, Paula A. Shaklee, Lief V. Gallagher, Thomas E. Munton, Holger Klinck, R.J. Gutiérrez, M. Zachariah. Peery. 2023. Estimating population size for California spotted owls and barred owls across the Sierra Nevada ecosystem with bioacoustics
• Kate A. McGinn, M. Zachariah Peery, Ceeanna J. Zulla, William J. Berigan, Zachary A. Wilkinson, Josh M. Barry, John J. Keane, Benjamin Zuckerberg. 2023. A climate-vulnerable species uses cooler forest microclimates during heat waves
• Ceeanna J. Zulla, H. Anu Kramer, Gavin M. Jones, John J. Keane, Kevin N. Roberts, Brian P. Dotters, Sarah Sawyer, Sheila A. Whitmore, William J. Berigan, Kevin G. Kelly, Amy K. Wray, M. Zachariah Peery. 2022. Large trees and forest heterogeneity facilitate prey capture by California Spotted Owls
• Dana S. Reid, Connor M. Wood, Sheila A. Whitmore, William J. Berigan, H. Anu Kramer, Nicholas F. Kryshak, John J. Keane, Sarah Sawyer, R. J. Gutiérrez, Holger Klinck, M. Zachariah. Peery. 2022. Breeding status shapes territoriality and vocalization patterns in spotted owls
• Nicholas F. Kryshak, Emily D. Fountain, Daniel F. Hofstadter, Brian P. Dotters, Kevin N. Roberts, Connor M. Wood, Kevin G. Kelly, Isabel F. Schwarcz, Paige J. Kulzer, Amy K. Wray, H. Anu Kramer, John P. Dumbacher, John J. Keane, Paula A. Shaklee, R.J. Gutiérrez, M. Zachariah. Peery. 2022. DNA metabarcoding reveals the threat of rapidly expanding barred owl populations to native wildlife in western North America
• Daniel Hofstadter, Nicholas Kryshak, Connor Wood, Brian Dotters, Kevin Roberts, Kevin Kelly, John J. Keane, Sarah Sawyer, Paula A. Shaklee, H Anu Kramer, RJ Gutiérrez, M Zachariah. Peery. 2022. Arresting the spread of invasive species in continental systems
• Gavin M. Jones, Alisa R. Keyser, A. Leroy Westerling, W. Jonathan Baldwin, John J. Keane, Sarah Sawyer, John D. J. Clare, R. J. Gutierrez, M. Zachariah Peery. 2021. Forest restoration limits megafires and supports species conservation under climate change
• Anu Kramer, Gavin M. Jones, Sheila A. Whitmore, John J. Keane, Fidelis A. Atuo, Brian P. Dotters, Sarah Sawyer, Sarah L. Stock, R.J. Gutierrez, M. Zachariah. Peery. 2021. California spotted owl habitat selection in a fire-managed landscape suggests conservation benefit of restoring historical fire regimes
• Daniel Hofstadter, Nicholas Kryshak, Mourad Gabriel, Connor Wood, Greta Wengert, Brian Dotters, Kevin Roberts, Emily Fountain, Kevin Kelly, John J. Keane, Sheila Whitmore, William Berigan, M Zachariah. Peery. 2021. High rates of anticoagulant rodenticide exposure in California Barred Owls are associated with the wildland–urban interface
• H. Anu Kramer, Gavin M. Jones, Van R. Kane, Bryce Bartl-Geller, Jonathan T. Kane, Sheila A. Whitmore, William J. Berigan, Brian P. Dotters, Kevin N. Roberts, Sarah Sawyer, John J. Keane, Malcolm P. North, R. J. Gutierrez, M. Zachariah Peery. 2021. Elevational gradients strongly mediate habitat selection patterns in a nocturnal predator
• Connor M. Wood, Nick Kryshak, Michaela Gustafson, Daniel F. Hofstadter, Brendan K. Hobart, Sheila A. Whitmore, Brian P. Dotters, Kevin N. Roberts, John J. Keane, Sarah Sawyer, Rocky J. Gutiérrez, M. Zachariah. Peery. 2021. Density dependence influences competition and hybridization at an invasion front
• Connor M. Wood, Ceeanna Zulla, Sheila Whitmore, Dana Reid, H. Anu Kramer, John J. Keane, Sarah Sawyer, Kevin N. Roberts, Brian P. Dotters, Holger Klinck, William Berigan, R. J. Gutiérrez, M. Zachariah. Peery. 2021. Illuminating the nocturnal habits of owls with emerging tagging technologies
• Brendan K. Hobart, H. Anu Kramer, Gavin M. Jones, Brian P. Dotters, Sheila A. Whitmore, John J. Keane, M. Zachariah. Peery. 2021. Stable isotopes reveal unexpected relationships between fire history and the diet of Spotted Owls
• Gavin M. Jones, R. J. Gutierrez, William M. Block, Peter C. Carlson, Emily J. Comfort, Samuel A. Cushman, Raymond J. Davis, Stephanie A. Eyes, Alan B. Franklin, Joseph L. Ganey, Shaula Hedwall, John J. Keane, Rodd Kelsey, Damon B. Lesmeister, Malcolm P. North, Susan L. Roberts, Jeremy T. Rockweit, Jamie S. Sanderlin, Sarah C. Sawyer, Ben Solvesky, Douglas J. Tempel, Ho Yiwan, A. Leroy Westerling, Gary C. White, M. Zachariah Peery. 2020. Spotted owls and forest fire: Comment
• Connor M. Wood, Holger Klinck, Michaela Gustafson, John J. Keane, Sarah C. Sawyer, R. J. Gutiérrez, M. Zachariah. Peery. 2020. Using the ecological significance of animal vocalizations to improve inference in acoustic monitoring programs
• Connor Wood, R.J. Gutiérrez, John J. Keane, M. Zachariah Peery. 2020. Early detection of rapid Barred Owl population growth within the range of the California Spotted Owl advises the Precautionary Principle
• Kevin Roberts, Sheila Whitmore, Brian P. Dotters, Fidelis A. Atuo, Martin G. Raphael, Sarah C. Sawyer, John J. Keane, R.J. Gutiérrez, M. Zachariah Peery. 2019. Resource selection by GPS-tagged California spotted owls in mixed-ownership forests
• Connor M. Wood, Viorel D. Popescu, Holger Klinck, John J. Keane, R.J. Gutiérrez, Sarah C. Sawyer, M. Zachariah. Peery. 2019. Detecting small changes in populations at landscape scales: a bioacoustic site-occupancy framework
• Brendan K. Hobart, Gavin M. Jones, Kevin N. Roberts, Brian P. Dotters, Sheila A. Whitmore, William J. Berigan, Martin G. Raphael, John J. Keane, R.J. Gutiérrez, M. Zachariah. Peery. 2019. Trophic interactions mediate the response of predator populations to habitat change
• Brendan K. Hobart, Kevin N. Roberts, Brian P. Dotters, William J. Berigan, Sheila A. Whitmore, Martin G. Raphael, John J. Keane, R.J. Gutiérrez, M. Zachariah. Peery. 2019. Site occupancy and reproductive dynamics of California spotted owls in a mixed-ownership landscape
• Scott Stephens, Leda Kobziar, Brandon Collins, Raymond Davis, Peter Fulé, William Gaines, Joseph L. Ganey, Jim M. Guldin, Paul F. Hessburg, Kevin Hiers, Serra J. Hoagland, John J. Keane, Ronald Masters, Ann McKellar, Warren Montague, Malcolm P. North, Thomas Spies. 2019. Is fire “for the birds”? How two rare species influence fire management across the US
• Zach Peery, David Wiens, Robin Bown, Peter C. Carlson, Katie Dugger, Jack Dumbacher, Alan B. Franklin, Keith A. Hamm, Mark Higley, John J. Keane. 2018. Barred owl research needs and prioritization in California
• Lief V. Gallagher, John J. Keane, Paula A. Shaklee, H. Anu Kramer, Ross A. Gerrard. 2018. Spotted owl foraging patterns following fuels treatments, Sierra Nevada, California
• Connor M. Wood, Sheila A. Whitmore, R.J. Gutiérrez, Sarah C. Sawyer, John J. Keane, M. Zachariah. Peery. 2018. Using metapopulation models to assess species conservation–ecosystem restoration trade-offs
• Gavin M. Jones, John J. Keane, R. J. Gutiérrez, M. Zachariah. Peery. 2018. Declining old-forest species as a legacy of large trees lost
• Malcolm P. North, Jonathan T. Kane, Van R. Kane, Gregory P. Asner, William Berigan, Derek J. Churchill, Scott Conway, R.J. Gutiérrez, Sean Jeronimo, John J. Keane, Alexander Koltunov, Tina Mark, Monika Moskal, Tom E. Munton, Zachary Peery, Carlos Ramirez, Rahel Sollmann, Angela M. White, Sheila Whitmore. 2017. Cover of tall trees best predicts California spotted owl habitat
• John J. Keane. 2017. Threats to the viability of California spotted owls
• Malcolm P. North, Mark W. Schwartz, Brandon M. Collins, John J. Keane. 2017. Current and projected condition of mid-elevation Sierra Nevada forests
• Brian S. Cade, Barry R. Noon, Rick D. Scherer, John J. Keane. 2017. Logistic quantile regression provides improved estimates for bounded avian counts: a case study of California Spotted Owl fledgling production
• Heather A. Kramer, Brandon M. Collins, Lief V. Gallagher, John J. Keane, Scott L. Stephens, Maggi Kelly. 2016. Accessible light detection and ranging: estimating large tree density for habitat identification
• Mary M. M. Conner, John J. Keane, Lief V. Gallagher, Tom E. Munton, Paula A. Shaklee. 2016. Comparing estimates of population change from occupancy and mark-recapture models for a territorial species
• Scott L. Stephens, Jay D. Miller, Brandon M. Collins, Malcolm P. North, John J. Keane, Susan L. Roberts. 2016. Wildfire impacts on California spotted owl nesting habitat in the Sierra Nevada
• Douglas J. Tempel, John J. Keane, R. J. Gutierrez, Jared Wolfe, Gavin M. Jones, Alexander Koltunov, Carlos M. Ramirez, William J. Berigan, Lief V. Gallagher, Tom E. Munton, Paula A. Shaklee, Sheila A. Whitmore, M. Zachariah Peery. 2016. Meta-analysis of California Spotted Owl (Strix occidentalis occidentalis) territory occupancy in the Sierra Nevada: habitat associations and their implications for forest management
• Paul F. Hessburg, Thomas Spies, David A. Perry, Carl N. Skinner, Alan H. Taylor, Peter M. Brown, Scott L. Stephens, Andrew J. Larson, Derek J. Churchill, Nicholas Povak, Peter Singleton, Brenda McComb, Bill Zielinski, Brandon M. Collins, R. Brion Salter, John J. Keane, Jerry F. Franklin, Greg Riegel. 2016. Tamm Review: Management of mixed-severity fire regime forests in Oregon, Washington, and Northern California
• Jennifer E. Carlson, Douglas D. Piirto, John J. Keane, Samantha J. Gill. 2015. Estimating site occupancy and detection probabilities for cooper's and sharp-shinned hawks in the Southern Sierra Nevada
• Joanna X. Wu, Rodney B. Siegel, Helen L. Loffland, Morgan W. Tingley, Sarah L. Stock, Kevin N. Roberts, John J. Keane, Joseph R. Medley, Roy Bridgman, Chris Stermer. 2015. Diversity of great gray owl nest sites and nesting habitats in California
• Scott L. Stephens, Seth W. Bigelow, Ryan D. Burnett, Brandon M. Collins, Lief V. Gallagher, John J. Keane, Douglas A. Kelt, Malcolm P. North, Lance J. Roberts, Peter Stine, Dirk H. Van Vuren. 2014. California spotted owl, songbird, and small mammal responses to landscape fuel treatments
• Joshua Hull, Andrew Englis, Joseph R. Medley, Eric P. Jepsen, James R. Duncan, Holly B. Ernest, John J. Keane. 2014. A new subspecies of great gray owl (Strix nebulosa) in the Sierra Nevada of California, U.S.A.
• John J. Keane. 2014. California spotted owl: scientific considerations for forest planning
• Malcolm P. North, Brandon M. Collins, John J. Keane, Jonathan W. Long, Carl N. Skinner, Bill Zielinski. 2014. Synopsis of emergent approaches
• Mary M. Conner, John J. Keane, Lief V. Gallagher, Gretchen A. Jehle, Tom E. Munton, Paula A. Shaklee, Ross A. Gerrard. 2013. Realized population change for long-term monitoring: California spotted owl case study
• J.M. Hull, A.M. Fish, John J. Keane, Sylvia Mori, B.J Sacks, A.C. Hull. 2010. Estimation of Species Identification Error: Implications for Raptor Migration Counts and Trend Estimation
• J.A. Blakesley, M.E. Seamans, M.M. Connor, A.B. Franklin, G.C. White, R.J. Gutierrez, J.E. Hines, J.D. Nichols, Tom E. Munton, D.W.H. Shaw, John J. Keane, G.N. Steger, T.L. McDonald. 2010. Population dynamics of the California spotted owl in the Sierra Nevada, California
• J.M. Hull, John J. Keane, W.K. Savage, S.A. Godwin, J. Shafer, E.P. Jepsen, R. Gerhardt, C. Stermer, H.B. Ernest. 2010. Range-wide genetic differentiation among North American great gray owls (Strix nebulosa) reveals a distinct lineage limited to the Sierra Nevada, California
• Patricia N. Manley, Bill Zielinski, C.M. Stuart, John J. Keane, A.J. Lind, C. Brown, B.L. Plymale, C.O. Napper. 2000. Monitoring ecosystems in the Sierra Nevada: the conceptual model foundation

• University of California, Ph.D., Ecology, 1999
• University of California, M.S., Wildland Resources Science, 1991
• San Francisco State University, B.S., Zoology, 1986