John R. Squires, PhD

Treesearch Publications

• Thomas W. Glass, Jeffery P. Copeland, Lucretia E. E. Olson, John S. Waller, John R. Squires. 2025. Wolverines use spatial memory to plan efficient routes through rugged terrain
• Heather Abernathy, Mark Ditmer, Seth B. Hibbard, Jason V Lombardi, Thomas R. Stephenson, John R. Squires, Jennifer N. Newton, Sarah R. Dewey, Katherine Zeller, Michael K. Schwartz, George Wittemyer. 2025. A case for human mobility data applications in wildlife management
• Sylvia Kantor, Katherine Zeller, Mark Ditmer, John R. Squires, Jesse Barber, James Wilder, Ashley Egan. 2025. How human-generated noise disrupts wildlife: The unseen impact of outdoor recreation
• Jacob S. Ivan, Karen E. Hodges, Joseph D. Holbrook, Ron A. Moen, Lucretia E. E. Olson, John R. Squires, Jennifer H. Vashon. 2024. Reply to Thornton and Murray: Models for Canada lynx conservation planning require nuance
• Casey C. Day, Erin L. Landguth, Michael A. Sawaya, Anthony P. Clevenger, Robert A. Long, Zachary Holden, Jocelyn R. Akins, Robert B. Anderson, Keith B. Aubry, Mirjam Barrueto, Nichole L. Bjornlie, Jeffrey P. Copeland, Jason T. Fisher, Anne Forshner, Justin A. Gude, Doris Hausleitner, Nichole A. Heim, Kimberly S. Heinemeyer, Anne Hubbs, Robert M. Inman, Scott Jackson, Michael Jokinen, Nathan P. Kluge, Andrea Kortello, Deborah L. Lacroix, Luke Lamar, Lisa I Larson, Jeffrey C. Lewis, Dave Lockman, Michael K. Lucid, Paula MacKay, Audrey J. Magoun, Michelle L. McLellan, Katie Moriarty, Cory E. Mosby, Garth Mowat, Clifford G. Nietvelt, David Paetkau, Eric C. Palm, Kylie J. S. Paul, Kristine L. Pilgrim, Catherine M. Raley, Michael K. Schwartz, Matthew A. Scrafford, John R. Squires, Zachary J. Walker, John S. Waller, Richard D. Weir, Katherine Zeller. 2024. Genetic connectivity of wolverines in western North America
• John R. Squires, Lucretia E. E. Olson, Jacob S. Ivan, Peter M. McDonald, Joseph D. Holbrook. 2024. Anthropogenically protected but naturally disturbed: A specialist carnivore at its southern range periphery
• Katherine Zeller, Mark Ditmer, John R. Squires, William L. Rice, James Wilder, Don DeLong, Ashley Egan, Niah Pennington, Chris A. Wang, Jacinta Plucinski, Jesse R. Barber. 2024. Experimental recreationist noise alters behavior and space use of wildlife
• Austin B. Smith, John R. Squires, Nichole L. Bjornlie, Joseph D. Holbrook. 2023. Divergent or convergent: How do forest carnivores use time in the Greater Yellowstone Ecosystem?
• John R. Squires, Jacob S. Ivan, Kelsey E. Paolini, Lucretia E. E. Olson, Gavin M. Jones, Joseph D. Holbrook. 2023. Keystone structures maintain forest function for Canada lynx after large-scale spruce beetle outbreak
• Simona Kraberger, Laurel E. K. Serieys, Seth P. D. Riley, Kara Schmidlin, Eric S. Newkirk, John R. Squires, Christopher B. Buck, Arvind Varsani. 2023. Novel polyomaviruses identified in fecal samples from four carnivore species
• Lucretia E. E. Olson, Justin S. Crotteau, John R. Squires. 2023. Compounding effects of wildfire and forest management on Canada lynx: The importance of management mosaics
• Lucretia E. E. Olson, Justin S. Crotteau, Shelagh Fox, Gary Hanvey, Joseph D. Holbrook, Scott Jackson, John R. Squires. 2023. Effects of compound disturbance on Canada lynx and snowshoe hare: Wildfire and forest management influence timing and intensity of use
• Josh McDaniel, Dave E. Calkin, Justin S. Crotteau, Mark A. Finney, Sam Larkin, Michael Johnston, Serra J. Hoagland, Dylan Johnson, Sarah McCaffrey, John R. Squires, Shawn P. Urbanski. 2022. Getting more fire on the ground: Landscape-scale prescribed burning supported by science
• Austin B. Smith, Jody C. Vogeler, Nichole L. Bjornlie, John R. Squires, Neal C. Swayze, Joseph D. Holbrook. 2022. Spaceborne LiDAR and animal-environment relationships: An assessment for forest carnivores and their prey in the Greater Yellowstone Ecosystem
• Andrea Watts, John R. Squires, Joseph Holbrook, Jake Ivan, Lucretia E. E. Olson, Peter McDonald, Kirby Self. 2021. Forest understories became the story: Lynx, bark beetle mortality, and salvage logging
• Rohan Antonio Bandoo, Joshua Bautista, Michael Lund, Eric Newkirk, John R. Squires, Arvind Varsani, Simona Kraberger. 2021. Identification of novel circovirus and anelloviruses from wolverines using a non-invasive faecal sampling approach
• Lucretia E. E. Olson, Nichole Bjornlie, Gary Hanvey, Joseph D. Holbrook, Jacob S. Ivan, Scott Jackson, Brian Kertson, Travis King, Michael Lucid, Dennis Murray, Robert Naney, John Rohrer, Arthur Scully, Daniel Thornton, Zachary Walker, John R. Squires. 2021. Improved prediction of Canada lynx distribution through regional model transferability and data efficiency
• Simona Kraberger, Laurel EK. Serieys, Cécile Richet, Nicholas M. Fountain-Jones, Guy Baele, Jacqueline M. Bishop, Mary Nehring, Jacob S. Ivan, Eric S. Newkirk, John R. Squires, Michael C. Lund, Seth PD. Riley, Christopher C. Wilmers, Paul D. van Helden, Koenraad Van Doorslaer, Melanie Culver, Sue VandeWoude, Darren P. Martin, Arvind Varsani. 2021. Complex evolutionary history of felid anelloviruses
• Niko Balkenhol, Michael K. Schwartz, Robert M. Inman, Jeffrey P. Copeland, John R. Squires, Neil J. Anderson, Lisette P. Waits. 2020. Landscape genetics of wolverines (Gulo gulo): Scale-dependent effects of bioclimatic, topographic, and anthropogenic variables
• John R. Squires, Lucretia E. E. Olson, Zachary P. Wallace, Robert J. Oakleaf, Patricia L. Kennedy. 2020. Resource selection of apex raptors: Implications for siting energy development in sagebrush and prairie ecosystems
• John R. Squires, Joseph D. Holbrook, Lucretia E. E. Olson, Jacob S. Ivan, Randal W. Ghormley, Rick L. Lawrence. 2020. A specialized forest carnivore navigates landscape-level disturbance: Canada lynx in spruce-beetle impacted forests
• John R. Squires, Lucretia E. E. Olson, Elizabeth K. Roberts, Jacob S. Ivan, Mark Hebblewhite. 2019. Winter recreation and Canada lynx: Reducing conflict through niche partitioning
• Sue Miller, Lucretia E. E. Olson, John R. Squires, Elizabeth Roberts, Jake Ivan. 2019. Winter sports and wildlife: Can Canada lynx and winter recreation share the same slope?
• Joseph D. Holbrook, John R. Squires, Barry Bollenbacher, Russ Graham, Lucretia E. E. Olson, Gary Hanvey, Scott Jackson, Rick L. Lawrence, Shannon L. Savage. 2019. Management of forests and forest carnivores: Relating landscape mosaics to habitat quality of Canada lynx at their range periphery
• Joseph D. Holbrook, Lucretia E. E. Olson, Nicholas J. DeCesare, Mark Hebblewhite, John R. Squires, Robin Steenweg. 2019. Functional responses in habitat selection: Clarifying hypotheses and interpretations
• Tracey N. Johnson, Kristen Nasman, Zachary P. Wallace, Lucretia E. E. Olson, John R. Squires, Ryan M. Nielson, Patricia L. Kennedy. 2019. Survey design for broad-scale, territory-based occupancy monitoring of a raptor: Ferruginous hawk (Buteo regalis) as a case study
• Kimberly Heinemeyer, John R. Squires, Mark Hebblewhite, Julia J. O’Keefe, Joseph D. Holbrook, Jeffrey Copeland. 2019. Wolverines in winter: Indirect habitat loss and functional responses to backcountry recreation
• Kevin S. McKelvey, Jessie Golding, Daniel H. Mason, Joseph Dysthe, Kristine L. Pilgrim, John R. Squires, Thomas W. Franklin, Keith B. Aubry, Robert A. Long, Samuel Greaves, Catherine M. Raley, Scott Jackson, Paula MacKay, Joshua Lisbon, Joel D. Sauder, Michael T. Pruss, Don Heffington, Michael K. Schwartz. 2019. Using environmental DNA methods to improve winter surveys for rare carnivores: DNA from snow and improved noninvasive techniques
• Jessie Golding, Michael K. Schwartz, Kevin S. McKelvey, John R. Squires, Scott D. Jackson, Cara Staab, Rema B. Sadak. 2018. Multispecies mesocarnivore monitoring: USDA Forest Service multiregional monitoring approach
• Lucretia E. E. Olson, John R. Squires, Elizabeth K. Roberts, Jacob S. Ivan, Mark Hebblewhite. 2018. Sharing the same slope: Behavioral responses of a threatened mesocarnivore to motorized and non-motorized winter recreation
• Joseph D. Holbrook, John R. Squires, Barry Bollenbacher, Russell T. Graham, Lucretia E. E. Olson, Gary Hanvey, Scott Jackson, Rick L. Lawrence. 2018. Spatio-temporal responses of Canada lynx (Lynx canadensis) to silvicultural treatments in the Northern Rockies, U.S.
• Shannon L. Savage, Rick L. Lawrence, John R. Squires. 2017. Mapping post-disturbance forest landscape composition with Landsat satellite imagery
• Lucretia E. E. Olson, John R. Squires, Robert J. Oakleaf, Zachary P. Wallace, Patricia L. Kennedy. 2017. Predicting above-ground density and distribution of small mammal prey species at large spatial scales
• Lucretia E. E. Olson, John R. Squires, Elizabeth K. Roberts, Aubrey D. Miller, Jacob S. Ivan, Mark Hebblewhite. 2017. Modeling large-scale winter recreation terrain selection with implications for recreation management and wildlife
• Aubrey Miller, Jerry J. Vaske, John R. Squires, Lucretia E. E. Olson. 2017. Does zoning winter recreationists reduce recreation conflict?
• Phillip E. Baigas, John R. Squires, Lucretia E. E. Olson, Jacob S. Ivan, Elizabeth. K. Roberts. 2017. Using environmental features to model highway crossing behavior of Canada lynx in the Southern Rocky Mountains
• Joseph D. Holbrook, John R. Squires, Lucretia E. E. Olson, Rick L. Lawrence, Shannon L. Savage. 2016. Multi-scale habitat relationships of snowshoe hares (Lepus americanus) in the mixed conifer landscape of the Northern Rockies, USA: Cross-scale effects of horizontal cover with implications for forest management
• Zachary P. Wallace, Patricia L. Kennedy, John R. Squires, Robert J. Oakleaf, Lucretia E. E. Olson, Katie M. Dugger. 2016. Re-occupancy of breeding territories by ferruginous hawks in Wyoming: Relationships to environmental and anthropogenic factors
• Zachary P. Wallace, Patricia L. Kennedy, John R. Squires, Lucretia E. E. Olson, Robert J. Oakleaf. 2016. Human-made structures, vegetation, and weather influence ferruginous hawk breeding performance
• Zachary A. Holden, Alan Swanson, Anna E. Klene, John T. Abatzoglou, Solomon Z. Dobrowski, Samuel A. Cushman, John R. Squires, Gretchen Moisen, Jared W. Oyler. 2016. Development of high-resolution (250 m) historical daily gridded air temperature data using reanalysis and distributed sensor networks for the US northern Rocky Mountains
• Lucretia E. E. Olson, Robert J. Oakleaf, John R. Squires, Zachary P. Wallace, Patricia L. Kennedy. 2015. Nesting pair density and abundance of ferruginous hawks (Buteo regalis) and golden eagles (Aquila chrysaetos) from aerial surveys in Wyoming
• Shannon L. Savage, Rick L. Lawrence, John R. Squires. 2015. Predicting relative species composition within mixed conifer forest pixels using zero‐inflated models and Landsat imagery
• Kevin S. McKelvey, Keith B. Aubry, Neil J. Anderson, Anthony P. Clevenger, Jeffrey P. Copeland, Kimberley S. Heinemeyer, Robert M. Inman, John R. Squires, John S. Waller, Kristine L. Pilgrim, Michael K. Schwartz. 2014. Recovery of wolverines in the western United States: Recent extirpation and recolonization or range retraction and expansion?
• John R. Squires, Nicholas J. DeCesare, Lucretia E. Olson, Jay A. Kolbe, Mark Hebblewhite, Sean A. Parks. 2013. Combining resource selection and movement behavior to predict corridors for Canada lynx at their southern range periphery
• Nathan D. Berg, Eric M. Gese, John R. Squires, Lise M. Aubry. 2012. Influence of forest structure on the abundance of snowshoe hares in western Wyoming
• John R. Squires, Nicholas J. DeCesare, Mark Hebblewhite, Joel Berger. 2012. Missing lynx and trophic cascades in food webs: A reply to Ripple et al.
• John R. Squires, Lucretia E. Olson, David Turner, Nicholas J. DeCesare, Jay A. Kolbe. 2012. Estimating detection probability for Canada lynx Lynx canadensis using snow-track surveys in the northern Rocky Mountains, Montana, USA
• Jeffrey P. Copeland, Kevin S. McKelvey, Michael K. Schwartz, Jeremy S. Littell, Keith B. Aubry, John R. Squires, Sean A. Parks, Marketa M. Elsner, Guillaume S. Mauger. 2011. Climate change predicted to shift wolverine distributions, connectivity, and dispersal corridors
• Lucretia E. Olson, John R. Squires, Nicholas J. DeCesare, Jay A. Kolbe. 2011. Den use and activity patterns in female Canada lynx (Lynx Canadensis) in the northern Rocky Mountains
• Kerry Fanson, Nadja C. Wielebnowski, Tanya M. Shenk, Walter J. Jakubas, John R. Squires, Jeffrey R. Lucas. 2010. Patterns of testicular activity in captive and wild Canada lynx (Lynx canadensis)
• Kerry V. Fanson, Nadja C. Wielebnowski, Tanya M. Shenk, Jennifer H. Vashon, John R. Squires, Jeffrey R. Lucas. 2010. Patterns of ovarian and luteal activity in captive and wild Canada lynx (Lynx canadensis)
• John R. Squires, Nicholas J. DeCesare, Jay A. Kolbe, Leonard F. Ruggiero. 2010. Seasonal resource selection of Canada lynx in managed forests of the northern Rocky Mountains
• Jeff Copeland, Kevin S. McKelvey, Keith B. Aubry, A. Landa, J. Persson, R. M. Inman, J. Krebs, E. Lofroth, H. Golden, John R. Squires, A. Magoun, Michael K. Schwartz, J. Wilmot, C. L. Copeland, Richard Yates, I. Kojola, R. May. 2010. The bioclimatic envelope of the wolverine (Gulo gulo): do climatic constraints limit its geographic distribution?
• Michael K. Schwartz, Jeffrey P. Copeland, Neil J. Anderson, John R. Squires, Robert M. Inman, Kevin S. McKelvey, Kristy L. Pilgrim, Lisette P. Waits, Samuel A. Cushman. 2009. Wolverine gene flow across a narrow climatic niche
• John R. Squires. 2008. Predicting the effects of forest management on lynx populations
• John R. Squires. 2008. The Goshawk - Robert Kenward [Book Review]
• John R. Squires, Nicholas J. Decesare, Jay A. Kolbe, Leonard F. Ruggiero. 2008. Hierarchical den selection of Canada lynx in western Montana
• Jay A. Kolbe, John R. Squires. 2007. Circadian activity patterns of Canada lynx in western Montana
• John R. Squires, Leonard F. Ruggiero. 2007. Winter prey selection of Canada lynx in northwestern Montana
• John R. Squires, Jeffrey P. Copeland, Todd J. Ulizio, Michael K. Schwartz, Leonard F. Ruggiero. 2007. Sources and patterns of wolverine mortality in western Montana
• Jay A. Kolbe, John R. Squires, Daniel H. Pletscher, Leonard F. Ruggiero. 2007. The effect of snowmobile trails on coyote movements within lynx home ranges
• Michael K. Schwartz, Keith B. Aubry, Kevin S. McKelvey, Kristine L. Pilgrim, Jeffrey P. Copeland, John R. Squires, Robert M. Inman, Samantha M. Wisely, Leonard F. Ruggiero. 2007. Inferring geographic isolation of wolverines in California using historical DNA
• Jay A. Kolbe, John R. Squires. 2006. A longevity record for Canada lynx, Lynx canadensis, in western Montana
• John R. Squires, Patricia L. Kennedy. 2006. Northern goshawk ecology: An assessment of current knowledge and information needs for conservation and management
• Todd J. Ulizio, John R. Squires, Daniel H. Pletscher, Michael K. Schwartz, James J. Claar, Leonard F. Ruggiero. 2006. The efficacy of obtaining genetic-based identifications from putative wolverine snow tracks
• Kevin S. McKelvey, Jeffrey von Kienast, Keith B. Aubry, Gary M. Koehler, Bejamin T. Maletzke, John R. Squires, Edward L. Lindquist, Steve Loch, Michael K. Schwartz. 2006. DNA analysis of hair and scat collected along snow tracks to document the presence of Canada Lynx.
• John R. Squires, Robert Oakleaf. 2005. Movements of a male Canada lynx crossing the greater Yellowstone Area, including highways
• Douglas A. Boyce, Patricia L. Kennedy, Paul Beier, Michael F. Ingraldi, Susie R. MacVean, Melissa S. Siders, John R. Squires, Brian Woodbridge. 2005. When are goshawks not there? Is a single visit enough to infer absence at occupied nest areas? Journal of Raptor Research
• Nicholas J. DeCesare, John R. Squires, Jay A. Kolbe. 2005. Effect of forest canopy on GPS-based movement data
• John R. Squires. 2005. Conservation challenges of managing lynx
• Jay A. Kolbe, John R. Squires. 2004. Long distance movement by a coyote within the Rocky Mountains
• John R. Squires, Kevin S. McKelvey, Leonard F. Ruggiero. 2004. A snow-tracking protocol used to delineate local lynx, Lynx canadensis, distributions
• Jay A. Kolbe, John R. Squires, Thomas W. Parker. 2003. An effective box trap for capturing lynx
• Roman Biek, Randall L. Zarnke, Colin Gillin, Margaret Wild, John R. Squires, Mary Poss. 2002. Serologic survey for viral and bacterial infections in western populations of Canada Lynx (Lynx canadensis)
• Gregory W. McDaniel, Kevin S. McKelvey, John R. Squires, Leonard F. Ruggiero. 2000. Efficacy of lures and hair snares to detect lynx
• John R. Squires. 2000. Food habits of Northern Goshawks nesting in south central Wyoming
• John R. Squires, Tom Laurion. 1999. Lynx home range and movements in Montana and Wyoming: Preliminary results [Chapter 11]
• Keith B. Aubry, Gary M. Koehler, John R. Squires. 1999. Ecology of Canada lynx in southern boreal forests [Chapter 13]
• Keith B. Aubry, Steven W. Buskirk, Gary M. Koehler, Charles J. Krebs, Kevin S. McKelvey, John R. Squires. 1999. The scientific basis for lynx conservation: Qualified insights [Chapter 16]
• Keith B. Aubry, Leonard F. Ruggiero, John R. Squires, Kevin S. McKelvey, Gary M. Koehler, Steven W. Buskirk, Charles J. Krebs. 1999. Conservation of lynx in the United States: A systematic approach to closing critical knowledge gaps [Chapter 17]
• Leonard F. Ruggiero, Keith B. Aubry, Steven W. Buskirk, Gary M. Koehler, Charles J. Krebs, Kevin S. McKelvey, John R. Squires. 1999. The scientific basis for lynx conservation: Can we get there from here? [Chapter 18]
• Steven W. Buskirk, Leonard F. Ruggiero, Keith B. Aubry, Dean E. Pearson, John R. Squires, Kevin S. McKelvey. 1999. Comparative ecology of lynx in North America [Chapter 14]
• Leonard F. Ruggiero, Keith B. Aubry, Steven W. Buskirk, Gary M. Koehler, Charles J. Krebs, Kevin S. McKelvey, John R. Squires. 1999. Ecology and conservation of lynx in the United States
• John R. Squires. 1998. Attempted kleptoparasitism of osprey by great blue herons
• John R. Squires, Stanley H. Anderson. 1997. Changes in trumpeter swan (Cygnus buccinator) activities from winter to spring in the greater Yellowstone area
• John R. Squires, Richard T. Reynolds. 1997. Northern goshawk (Accipiter gentilis)
• John R. Squires, Leonard F. Ruggiero. 1996. Nest-site preference of northern goshawks in southcentral Wyoming
• John R. Squires. 1995. Carrion use by northern goshawks
• John R. Squires, Leonard F. Ruggiero. 1995. Winter movements of adult northern goshawks that nested in southcentral Wyoming
• John R. Squires, Stanley H. Anderson. 1995. Trumpeter swan food habitats in the greater Yellowstone ecosystem
• Gregory D. Hayward, John R. Squires. 1994. Viability analysis in biological evaluations: Concepts of population viability analysis, biological population, and ecological scale
• John R. Squires, Stanley H. Anderson, Robert Oakleaf. 1993. Home range size and habitat-use pattern of nesting prairie falcons near oil developments in northeastern Wyoming
• John R. Squires, Stanley H. Anderson, Robert Oakleaf. 1991. Prairie falcons quit nesting in response to spring snowstorm
• John R. Squires, Stanley H. Anderson, Robert Oakleaf. 1989. Food habits of nesting prairie falcons in Campbell County

• Project
  

  Wildlife responses to recreation noise

  https://research.fs.usda.gov/rmrs/projects/wildliferecnoise
  Station

  Rocky Mountain Research Station
• Program
  

  Wildlife Ecology

  https://research.fs.usda.gov/rmrs/programs/wildlife-ecology
  Station

  Rocky Mountain Research Station
• Project
  

  The Canada Lynx of the Rockies Research Program

  https://research.fs.usda.gov/rmrs/projects/lynxresearchprogram
  Station

  Rocky Mountain Research Station
• Project
  

  Planning a multiregional monitoring effort for forest carnivores

  https://research.fs.usda.gov/rmrs/projects/monitoringforestcarnivores
  Station

  Rocky Mountain Research Station

Multimedia

• Webinars

  Fire Science You Can Use: Emerging Insights in Fire Ecology

  https://research.fs.usda.gov/rmrs/products/multimedia/webinars/fire-science-you-can-use-emerging-insights-fire-ecology
• Video

  Meet Research Wildlife Biologist John Squires

  https://research.fs.usda.gov/rmrs/products/multimedia/videos/meet-research-wildlife-biologist-john-squires

• Colorado State University, B.S., Wildlife Biology, 1979
• University of Wyoming, M.S., Zoology, 1986
• University of Wyoming, Ph.D., Zoology, 1991