John Willis, PhD
Treesearch Publications
- Benjamin O. Knapp, Jacob S. Fraser, Joanne Baggs, Vikram Chhatre, David Clabo, Connor D. Crouch, Daniel C. C. Dey, Shaik M. Hossain, Steven B. Jack, John M. Kabrick, Kathryn R. Kidd, Steve McNulty, Carl Nordman, Matthew G. Olson, Kyle Palmquist, Matthew Pelkki, Christopher Schalk, Michael C. Stambaugh, J. Morgan Varner, John Willis. 2026. Shortleaf Pine (Pinus echinata): Management Challenges, Knowledge Gaps, and Opportunities for an Unsung Species
- John Willis, Christopher Schalk, Josh B. Pierce. 2025. Seed removal patterns of fire-sensitive tree species on floodplain and upland sites in the absence of fire
- Xiongwen Chen, John Willis, David C. Clabo. 2025. Longleaf pine growth divergence increases over time across its geographic range
- Xiongwen Chen, John Willis. 2025. Characterizing the growth of Pinus palustris and the relationship with cone production at the individual tree level
- John Willis, Ajay Sharma, Timothy M. Shearman, J. Morgan Varner, Justin McKeithen. 2025. Longleaf and shortleaf pine seedling fire tolerance is more sensitive to shade than encroaching hardwood species
- Ashton Shell, Ajay Sharma, John Willis, John Tracy, Adam Polinko, Santosh Ojha, Jason Vogel. 2025. Growth dynamics of longleaf pine during conversion to uneven-aged stands
- Michael Ulyshen, Cory K. Adams, Jacqui Adams, Susan B. Adams, Mickey Bland, Don C. Bragg, Chuck Burdine, Mac A. Callaham, Richard Chaney, Gregg Chapman, Patsy Clinton, Cinnamon Dixon, Jacob Floyd, Phillip Jordan, Tara L. Keyser, Stephanie H. Laseter, Rachel Mallinger, Virginia McDaniel, Bryan Mudder, C. Dana Nelson, Katherine Odanaka, A. Christopher Oishi, Shawna L. Reid, Samm Reynolds, Kevin Robertson, Daniel Saenz, Mary Anne Sayer, Nathan Schiff, Brian Scholtens, Joel Scott, Martin Spetich, Melanie K. Taylor, John Willis, Andrew D. Young. 2024. Spatiotemporal patterns of forest pollinator diversity across the southeastern United States
- Monika Rawat, Tamara Milton, Rachel Nation, John Willis, Stephen Brewer, Callie Schweitzer, Heather D. Alexander. 2024. Exploring Leaf Litter Flammability in Eastern U.S. Oak and Pine Forests: Implications for Fire Ecology
- Arthur Lamounier Moura, Heather D. Alexander, John Willis, Zhaofei Fan, Michael J. Aspinwall, Callie Schweitzer, Kevin M. Robertson. 2024. Exploring the Effects of Fire Return Intervals on Resource-Use Strategies of Pyrophytic and Mesophytic Species
- Heather D. Alexander, Josue Chevez Sahona, Brice Hanberry, Tamara Milton, Arthur Lamounier Moura, Monika Rawat, Courtney Siegert, John Willis. 2024. Mesophication of Oak Landscapes: Mechanisms and Management
- Robert Tatina, Brice Hanberry, John Willis. 2024. Differentiating historical open forests and current closed forests of the Coastal Plain, southeastern USA
- John Willis, Tamara F. Milton, Heather D. Alexander. 2024. Cone and fruit impacts on understory flammability depend on traits and forest floor coverage
- Jeffery B. Cannon, Brandon T. Rutledge, Joshua J. Puhlick, John Willis, Dale Brockway. 2024. Hurricanes Stimulate Cone Production in Longleaf Pine Woodlands
- Joshua J. Puhlick, Benjamin O. Knapp, Ian Goldberg, Brandon T. Rutledge, R. Scott Taylor, John Willis. 2024. Silvicultural Strategies for Promoting Longleaf Pine Recruitment in Multiaged Stands
- Edward Andrews, John Willis. 2024. Prescribed Burning has Little Effect on Longleaf Pine Wood Quality
- John Willis, Don C. Bragg, Jeffery B. Cannon, Kamal J.K. Gandhi, Kathryn R. Kidd, Adam D. Polinko, Joshua J. Puhlick, Dan Saenz, Mary Anne Sayer, Christopher Schalk, Andrew B. Self, Courtney M. Siegert, J. Morgan. Varner. 2024. Does Restoring Woodland Structure with Loblolly or Slash Pine Produce Desired Ecosystem Functions?
- Rachel E. Nation, Heather D. Alexander, John Willis, Michael Aspinwall, Jeffery B. Cannon. 2024. Does Longleaf Pine Cone Production Correlate with Seed Production, Size, and Germination Rate?
- John Willis, A.B. Self. 2024. Early silvicultural intervention affects species dominance in efforts to restore pine-hardwood mixedwoods following stand-replacing disturbance
- John Willis, Don C. Bragg, Jeffery B. Cannon, Kamal J. K. Gandhi, Kathryn R. Kidd, Adam D. Polinko, Joshua J. Puhlick, Daniel Saenz, Mary Anne Sword Sayer, Christopher Schalk, Andrew B. Self, Courtney M. Siegert, J. Morgan Varner. 2024. Assessing the potential impact of retaining native off‐site tree species in woodland restoration
- Jeffery B. Cannon, Brandon T. Rutledge, Joshua J. Puhlick, John Willis, Dale Brockway. 2024. Tropical cyclone winds and precipitation stimulate cone production in the masting species longleaf pine (Pinus palustris)
- Xiongwen Chen, John Willis, Qinfeng Guo. 2023. Patterns of Water Consumption in Longleaf Pine Restoration Areas and the Relationship with Cone Production
- Steven Cabrera, Heather D. Alexander, John Willis, Christopher J. Anderson. 2023. Midstory removal of encroaching species has minimal impacts on fuels and fire behavior regardless of burn season in a degraded pine-oak mixture
- Darcey Collins, Joshua J. Granger, Stephen Dicke, Krishna P. Poudel, Adam Polinko, John Willis. 2023. Crown lifting of low-density loblolly pine (Pinus taeda) plantations on nonindustrial private forest lands
- Xiongwen Chen, John Willis. 2023. Characterizing the change of annual cone production in longleaf pine forests
- Xiongwen Chen, John Willis. 2023. Individuals’ behaviors of cone production in longleaf pine trees
- John Willis, H.J. Renninger, D.K. Schnake, H.D. Alexander. 2023. Xerophytic hardwood retention promotes competition over facilitation in longleaf pine woodlands in the absence of fire
- Xiongwen Chen, Kimberly A. Bowman, John Willis. 2023. Critical slowing down in cone production of longleaf pine trees
- John Willis, Andrew B. Self, Courtney M. Siegert. 2022. Proceedings of the 21st Biennial Southern Silvicultural Research Conference
- Ashton B. Shell, Ajay Sharma, Jason G. Vogel, John Willis. 2022. Converting plantations to uneven-aged stands: effects of harvest type, group opening size, orientation, and canopy position on residual tree development
- Xiongwen Chen, John Willis. 2022. Interactions of biosphere and atmosphere within longleaf pine restoration areas
- John Willis, Michael A. Blazier. 2022. Competition intensity varies with hardwood species identity and constrains stand-level productivity in southeastern pine–hardwood mixtures compared to loblolly pine monocultures
- A.D. Polinko, John Willis, A. Sharma, J.M. Guldin. 2022. Stand-level structural characteristics dictate hurricane resistance and resilience more than silvicultural regime in longleaf pine woodlands
- John Willis, Ajay Sharma, John S. Kush. 2022. Seasonal biennial burning has negligible effects on longleaf pine basal area growth
- Andrew B. Self, John Willis. 2022. Eleven-year effects of mechanical site preparation on oaks planted on former agricultural fields
- Arun Regmi, Donald L. Grebner, John Willis, Robert K. Grala. 2022. Sawmill willingness to pay price premiums for higher quality pine sawtimber in the southeastern United States
- Xiongwen Chen, John Willis, Kimberly A. Bowman. 2022. Assessing the influence of climate on cone production of longleaf pine forests
- E.K. Babl-Plauche, H.D. Alexander, C.M. Siegert, John Willis, A.I. Berry. 2022. Mesophication of upland oak forests: Implications of species-specific differences in leaf litter decomposition rates and fuelbed composition
- Andrew Hacket‐Pain, Jessie J. Foest, Ian S. Pearse, Jalene M. LaMontagne, Walter D. Koenig, Giorgio Vacchiano, Michał Bogdziewicz, Thomas Caignard, Paulina Celebias, Joep Dormolen, Marcos Fernández‐Martínez, Jose V Moris, Ciprian Palaghianu, Mario Pesendorfer, Akiko Satake, Eliane Schermer, Andrew J. Tanentzap, Peter A. Thomas, Davide Vecchio, Andreas P. Wion, Thomas Wohlgemuth, Tingting Xue, Katharine Abernethy, Marie‐Claire Aravena Acuña, Marcelo Daniel Barrera, Jessica H. Barton, Stan Boutin, Emma R. Bush, Sergio Donoso Calderón, Felipe S. Carevic, Carolina Volkmer Castilho, Juan Manuel Cellini, Colin A. Chapman, Hazel Chapman, Francesco Chianucci, Patricia Costa, Luc Croisé, Andrea Cutini, Ben Dantzer, R. Justin DeRose, Jean‐Thoussaint Dikangadissi, Edmond Dimoto, Fernanda Lopes Fonseca, Leonardo Gallo, Georg Gratzer, David F. Greene, Martín A. Hadad, Alejandro Huertas Herrera, Kathryn J. Jeffery, Jill F. Johnstone, Urs Kalbitzer, Władysław Kantorowicz, Christie A. Klimas, Jonathan G. A. Lageard, Jeffrey Lane, Katharina Lapin, Mateusz Ledwoń, Abigail C. Leeper, Maria Vanessa Lencinas, Ana Cláudia Lira‐Guedes, Michael C. Lordon, Paula Marchelli, Shealyn Marino, Harald Schmidt Van Marle, Andrew G. McAdam, Ludovic R. W. Momont, Manuel Nicolas, Lúcia Helena Oliveira Wadt, Parisa Panahi, Guillermo Martínez Pastur, Thomas Patterson, Pablo Luis Peri, Łukasz Piechnik, Mehdi Pourhashemi, Claudia Espinoza Quezada, Fidel A. Roig, Karen Peña Rojas, Yamina Micaela Rosas, Silvio Schueler, Barbara Seget, Rosina Soler, Michael A. Steele, Mónica Toro‐Manríquez, Caroline E. G. Tutin, Tharcisse Ukizintambara, Lee White, Biplang Yadok, John Willis, Anita Zolles, Magdalena Żywiec, Davide Ascoli. 2022. MASTREE+: Time‐series of plant reproductive effort from six continents
- Arun Regmi, Donald Grebner, John Willis, Robert K. Grala. 2022. Price premium requirements for growing higher quality pine sawtimber in even-aged systems in the southeastern United States
- John Willis, Ajay Sharma, John S. Kush. 2021. Seasonality of biennial burning has no adverse effects on mature longleaf pine survival or productivity
- John Willis, David K. Schnake, Christopher S. DePerno, Marcus A. Lashley, Branson Wetzstein, Justin Yow. 2021. Tree encroachment impacts on seed predator selection and seedling establishment in degraded pine woodlands
- John Willis, Michael B. Walters, Evan J. Farinosi. 2021. Can coppicing planted saplings improve the growing position of mid-tolerant northern hardwood tree species in harvest gaps?
- David K. Schnake, Scott D. Roberts, John Willis, John D. Kushla, Ian A. Munn. 2021. Overstory retention and stock type impact survival and growth of underplanted shortleaf pine beneath a hardwood canopy
- John Willis, Constance A. Harrington, Leslie C. Brodie, Scott D. Roberts. 2021. Variable-density thinning promotes differential recruitment and development of shade tolerant conifer species after 17 years
- David Clabo, Jason Gordon, Arnold Brodbeck, John Willis, Shaun Tanger, Andrew Self. 2020. Managing mixed stands for southern forest landowners: three family stories for decision-making assistance
- Jason Gordon, Marc Measells, John Willis, Brady Self. 2020. Experiences from a forestry extension long-term impact evaluation
- Jason S. Gordon, John Willis, Robert K. Grala. 2020. Public and forest landowner attitudes towards longleaf pine ecosystem restoration using prescribed fire
- Andrew B. Self, John Willis, Andrew W. Ezell. 2020. Oak regeneration success in operational shelterwood cuts: year two results
- Jacob Henry, John Willis, David K. Schnake, Scott D. Roberts, Heather D. Alexander, Heidi Renninger. 2020. Early impacts of midstory hardwoods and overstory density on longleaf pine seedling establishment on xeric sites
- Arun Regmi, Donald L. Grebner, Robert K. Grala, John Willis. 2020. Preliminary evaluation of price premium required for growing higher quality loblolly and slash pines on extended rotation ages
- Michael B. Walters, Evan J. Farinosi, John Willis. 2020. Deer browsing and shrub competition set sapling recruitment height and interact with light to shape recruitment niches for temperate forest tree species
- Jason Gordon, Marc Measells, John Willis, Brady Self. 2020. Comparing social media and postal mailings in forestry extension program marketing
- Emily Babl, Heather D. Alexander, Courtney M. Siegert, John Willis. 2020. Could canopy, bark, and leaf litter traits of encroaching non-oak species influence future flammability of upland oak forests?
- John Willis, Jason S. Gordon, Shaun Tanger, Michael A. Blazier, Andrew B. Self, Arnold Brodbeck. 2019. Managing mixed stands: reassessing a forgotten stand type in the southeastern United States
- Emily K. Babl, Heather D. Alexander, Courtney M. Siegert, John Willis, Andrew I Berry. 2019. Species-specific mechanisms contributing to the mesophication of upland oak stands in the absence of fire
Citations of Non-Forest Service Publications
- -Willis, J.L., Schnake, D.K., Wetzstein, B., Yow, J., Guinto, D., Ulrich, S., DePerno, C.S., Lashley, M.A. (2019). Seed depredation negates the benefits of midstory hardwood removal on longleaf pine seedling establishment. Restoration Ecology. 0, 1-9
-Willis, J.L., Walters, M.B. (2018). Nutrition and mycorrhizae affect interspecific patterns of seedling growth on coarse wood and mineral soil substrates. Ecosphere. 9, 1-17
-Webster, C.R., Dickinson, Y.L. Burton, J.I, Frelich, L.E., Jenkins, M.A., Kern, C.C., Raymond, P., Saunders, M.R., Walters, M.B., Willis, J.L. (2018). Promoting and maintaining diversity in contemporary hardwood forests: confronting contemporary drivers of change and the loss of ecological memory. Forest Ecology and Management. 421, 98-108
-Willis, J.L., Roberts, S.D., Harrington, C.A. (2018). Variable density thinning promotes variable structural responses 14 years after treatment in the Pacific Northwest. Forest Ecology and Management. 410, 114-125
-Kern, C.C., Burton, J.I., Raymond, P., D’Amato, A.W., Keeton, W.S., Royo, A.A., Walters, M.B., Willis, J.L. (2017). Challenges facing gap-based silviculture and possible solutions for mesic northern forests in North America. Forestry. 89, 1-14
-Walters, M.B., Farinosi, E., Willis, J.L. (2016). Managing for diversity: harvest gap size drives complex light, vegetation, and deer herbivory impacts on tree seedlings. Ecosphere. 7, 1-29
-Willis, J.L, Walters, M.B. (2016). Local seed source availability limits young seedling populations for some species more than other factors in northern hardwood forests. Forest Science. 62, 440-448
-Willis, J.L., Walters, M.B., Gottschalk, K.W. (2015). Scarification and harvest gap size have interacting effects on young seedling establishment. Forest Ecology and Management. 347, 237-246
-Walters, M.B., Willis, J. L, Gottschalk, K.W. (2014). Can light and nitrate vs. ammonium growth responses among mixed hardwood-conifer forest species be predicted by their ecologies and help explain community scale tree diversity? Canadian Journal of Forest Research. 44, 1356-1368
- Michigan State University, Ph.D. Forestry and Evolutionary Ecology and Behavior Biology, Forest Ecology and Silviculture, 2014
- Colorado State University, M.S. Forest Science, Silviculture, 2009
- Colorado State University, B.S Forestry, Fire Science, 2007
Last updated March 10, 2026