Justin B. Runyon, PhD

Research Entomologist

Rocky Mountain Research Station

justin.runyon@usda.gov

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Treesearch Publications

Katherine Sparks, Sean L. Hoy-Skubik, Franklin Alongi, Justin B. Runyon, Katharine M. Banner, Brian V Smithers, Danielle E. M. Ulrich. 2025. Comparing juvenile physiology and morphology of two high-elevation pines, Pinus albicaulis and P. balfouriana
Thomas Kolb, Monica Gaylord, Amanda Grady, Matt Hansen, Ann M. Lynch, Joel McMillin, Justin B. Runyon. 2025. Does spruce aphid defoliation alter Engelmann spruce resistance to spruce beetle?
Sue Miller, Anthony D. Vaudo, Bryce A. Richardson, Sarah C. Barga, Justin B. Runyon. 2025. Bee nourished: How pollen nutrition shapes bee foraging habits and what it means for restoration
Justin B. Runyon. 2024. Revision of the genus Calyxochaetus Bigot (Diptera: Dolichopodidae: Sympycninae)
Will R. Glenny, Justin B. Runyon, Laura A. Burkle. 2024. Bumble bee diet breadth increases with local abundance and phenophase duration, not intraspecific variation in body size
Michael Ulyshen, Kimberly Ballare, Christopher J. Fettig, James W. Rivers, Justin B. Runyon. 2024. The value of forests to pollinating insects varies with forest structure, composition, and age
Justin B. Runyon, Will Glenny, Rebecca Jones. 2024. What do bees need? Understanding food and nesting sites in an ecotone
The Herbivory Variability Network, Justin B. Runyon. 2023. Plant size, latitude, and phylogeny explain within population variability in herbivory
Matheus M. M. Soares, Justin B. Runyon, Renato S. Capellari, Rosaly Alerocha. 2023. Review of Paraclius arcuatus species-group (Diptera, Dolichopodidae) with description of two new species
Will Glenny, Justin B. Runyon, Laura A. Burkle. 2023. Habitat characteristics structuring bee communities in a forest-shrubland ecotone
Kevin M. O’Neill, Ruth P. O’Neill, Casey M. Delphia, Laura A. Burkle, Justin B. Runyon. 2023. Diversity and distribution of orchid bees (Hymenoptera: Apidae, Euglossini) in Belize
Sylvia Kantor, Justin B. Runyon, Will Glenny, Laura Burkle, Jessie Salix, Don DeLong. 2023. Of bees and blooms: A new scorecard for selecting pollinator-friendly plants in restoration
Will Glenny, Justin B. Runyon, Laura A. Burkle. 2023. Plant selection for pollinator restoration in seminatural ecosystems
Justin B. Runyon, Eric Beckendorf, Louis S. Hesler. 2022. A new species of Chrysotus Meigen (Diptera, Dolichopodidae from soybean fields in South Dakota, USA
Justin B. Runyon, Barbara J. Bentz, Claire A. Qubain. 2022. Constitutive and induced defenses in long-lived pines do not trade off but are influenced by climate
David N. Soderberg, Barbara J. Bentz, Justin B. Runyon, Sharon M. Hood, Karen E. Mock. 2022. Chemical defense strategies, induction timing, growth, and trade-offs in Pinus aristata and Pinus flexilis
Shealyn C. Malone, Fabian D. Menalled, David K. Weaver, Tim F. Seipel, Megan L. Hofland, Justin B. Runyon, Maryse Bourgault, Darrin L. Boss, Amy M. Trowbridge. 2022. Cropping systems alter plant volatile emissions in the field through soil legacy effects
Christopher J. Fettig, Justin B. Runyon, Crystal S. Homicz, Patrick M. A. James, Michael Ulyshen. 2022. Fire and Insect Interactions in North American Forests
Christopher J. Fettig, A. Steven Munson, Justin B. Runyon, Leif A. Mortenson, Brytten E. Steed, Kenneth E. Gibson, Carl L. Jørgensen, Stephen Mckelvey, Joel D. McMillin, Jackson Audley, Jose F. Negrón. 2022. Ecological consequences of mountain pine beetle outbreaks in the Intermountain West
Will Glenny, Justin B. Runyon, Laura A. Burkle. 2022. A review of management actions on insect pollinators on public lands in the United States
Justin B. Runyon. 2022. Nematode-induced demasculinization of Nearctic Dolichopodidae (Diptera) with five new synonyms
Will Glenny, Justin B. Runyon, Laura Burkle. 2022. Assessing pollinator friendliness of plants and designing mixes to restore habitat for bees
George Poinar, Justin B. Runyon. 2021. Parasitylenchus myiophagus n. sp. (Nematoda: Parasitylenchidae), a tylenchid nematode parasite of long-legged flies (Diptera: Dolichopodidae)
Barbara J. Bentz, David Soderberg, Justin B. Runyon, Sharon M. Hood, Jim C. Vandygriff, Matt Hansen, Sky Stephens. 2021. Chapter 10 - Vulnerability of Rocky Mountain bristlecone pine to mountain pine beetle
Gabriel Foote, Justin B. Runyon, Christopher J. Fettig, Nathaniel Foote, Darrell W. Ross. 2021. Wild bee response to application of the Douglas-fir beetle anti-aggregation pheromone, 3-methylcyclohex-2-en-1-one
Justin B. Runyon. 2021. Invasive Species Science Update (No. 13)
Christopher J. Fettig, Sharon M. Hood, Justin B. Runyon, Chris M. Stalling. 2021. Bark beetle and fire interactions in western coniferous forests: Research findings
Therese M. M. Poland, Jennifer Juzwik, Allen Rowley, Cynthia Huebner, John Kilgo, Vanessa M. Lopez, Deanna H. Olson, Dean E. Pearson, Robert Progar, Robert Rabaglia, John Rothlisberger, Justin B. Runyon, Sharlene E. Sing. 2021. Management of Landscapes for Established Invasive Species
Deborah M. Finch, Jack L. Butler, Justin B. Runyon, Christopher J. Fettig, Francis F. Kilkenny, Shibu Jose, Susan J. Frankel, Samuel A. Cushman, Richard C. Cobb, Jeffrey S. Dukes, Jeffrey A. Hicke, Sybill K. Amelon. 2021. Effects of Climate Change on Invasive Species
Jackson Audley, Christopher J. Fettig, A. Steven Munson, Justin B. Runyon, Leif A. Mortenson, Brytten E. Steed, Kenneth E. Gibson, Carl L. Jørgensen, Stephen R. McKelvey, Joel D. McMillin, Jose F. Negrón. 2021. Dynamics of beetle-killed snags following mountain pine beetle outbreaks in lodgepole pine forests
John F. Gaskin, Erin Espeland, Casey D. Johnson, Diane L. Larson, Jane M. Mangold, Rachel A. McGee, Chuck Milner, Shishir Paudel, Dean E. Pearson, Lora B. Perkins, Chadley W. Prosser, Justin B. Runyon, Sharlene E. Sing, Zachary A. Sylvain, Amy J. Symstad, Daniel R. Tekiela. 2021. Managing invasive plants on Great Plains grasslands: A discussion of current challenges
Shealyn C. Malone, David K. Weaver, Tim F. Seipel, Fabian D. Menalled, Megan L. Hofland, Justin B. Runyon, Amy M. Trowbridge. 2020. Herbivore-induced volatile emissions are altered by soil legacy effects in cereal cropping systems
Justin B. Runyon, Christopher J. Fettig, Jared Trilling, A. Steven Munson, Leif A. Mortenson, Brytten E. Steed, Kenneth E. Gibson, Carl L. Jørgensen, Stephen Mckelvey, Joel D. McMillin, Jackson Audley, Jose Negron. 2020. Changes in understory vegetation including invasive weeds following mountain pine beetle outbreaks
Gabriel Foote, Nathaniel Foote, Justin B. Runyon, Darrell Ross, Christopher J. Fettig. 2020. Changes in the summer wild bee community following a bark beetle outbreak in a Douglas-fir forest
Laura A. Burkle, William R. Glenny, Justin B. Runyon. 2020. Intraspecific and interspecific variation in floral volatiles over time
Justin B. Runyon. 2020. The Dolichopodidae (Diptera) of Montserrat, West Indies
Christopher J. Fettig, Daniel S. Ott, A. Steven Munson, Justin B. Runyon, Darrell W. Ross. 2020. Physical and chemical characteristics of blue and Engelmann spruce relative to spruce beetle host selection and colonization
Jackson P. Audley, Christopher J. Fettig, A. Steven Munson, Justin B. Runyon, Leif Mortenson, Brytten E. Steed, Kenneth E. Gibson, Carl L. Jørgensen, Stephen Mckelvey, Joel D. McMillin, Jose Negron. 2020. Impacts of mountain pine beetle outbreaks on lodgepole pine forests in the Intermountain West, U.S., 2004–2019
Gabriel Foote, Christopher J. Fettig, Darrell Ross, Justin B. Runyon, Tom Coleman, Monica Gaylord, Andrew Graves, Joel McMillin, Leif Mortenson, Agenor Mafra-Neto. 2020. A biodegradable formulation of MCH (3-methylcyclohex-2-en-1-one) for protecting Pseudotsuga menziesii from Dendroctonus pseudotsugae (Coleoptera: Curculionidae) colonization
Justin B. Runyon. 2020. Invasive Species Science Update (No. 12)
Justin B. Runyon, Curtis A. Gray, Michael J. Jenkins. 2020. Volatiles of high-elevation five-needle pines: Chemical signatures through ratios and insight into insect and pathogen resistance
Casey M. Delphia, Justin B. Runyon, Kevin M. O’Neill. 2019. Clear plastic bags of bark mulch trap and kill female Megachile (Hymenoptera: Megachilidae) searching for nesting sites
Laura A. Burkle, Justin B. Runyon. 2019. Floral volatiles structure plant-pollinator interactions in a diverse community across the growing season
Curtis A. Gray, Justin B. Runyon, Michael J. Jenkins. 2019. Great Basin bristlecone pine volatiles as a climate change signal across environmental gradients
Justin B. Runyon. 2019. Two new species of Hurleyella Runyon and Robinson (Diptera: Dolichopodidae), with the first record from the Neotropics
Justin B. Runyon, M. Pollet. 2019. The genus Enlinia Aldrich in Chile (Diptera: Dolichopodidae), with the description of four new species
Justin B. Runyon. 2019. Invasive Species Science Update (No. 11)
Justin B. Runyon, Marc Pollet. 2018. Enlinia Aldrich, 1933 of Mitaraka, French Guiana (Diptera: Dolichopodidae)
Dean E. Pearson, Yvette K. Ortega, Justin B. Runyon, Jack L. Butler. 2018. Secondary invasion re-redefined: The distinction between invader-facilitated and invader-contingent invasions as subclasses of secondary invasion
William R. Glenny, Justin B. Runyon, Laura A. Burkle. 2018. Drought and increased CO2 alter floral visual and olfactory traits with context-dependent effects on pollinator visitation
Justin B. Runyon, Renato S. Capellari. 2018. Palpi aplenty: New species in the Chrysotus longipalpus species group (Diptera: Dolichopodidae)
Justin B. Runyon. 2018. Invasive Species Science Update (No. 10)
Art Borkent, Brian V. Brown, Peter H. Adler, Dalton de Souza Amorim, Kevin Barber, Daniel Bickel, Stephanie Boucher, Scott E. Brooks, John Burger, Zelia L. Burington, Renato S. Capellari, Daniel N. R. Costa, Jeffrey M. Cumming, Greg Curler, Carl W. Dick, John H. Epler, Eric Fisher, Stephen D. Gaimari, Jon Gelhaus, David A. Grimaldi, John Hash, Martin Hauser, Heikki Hippa, Sergio Ibanez-Bernal, Mathias Jaschhof, Elena P. Kameneva, Peter H. Kerr, Valery Korneyev, Cheslavo A. Korytkowski, Giar-Ann Kung, Gunnar Mikalsen Kvifte, Owen Lonsdale, Stephen A. Marshall, Wayne N. Mathis, Verner Michelsen, Stefan Naglis, Allen L. Norrbom, Steven Paiero, Thomas Pape, Alessandre Pereira-Colavite, Marc Pollet, Sabrina Rochefort, Alessandra Rung, Justin B. Runyon, Jade Savage, Vera C. Silva, Bradley J. Sinclair, Jeffrey H. Skevington, John O. Stireman, John Swann, F. Christian Thompson, Pekka Vilkamaa, Terry Wheeler, Terry Whitworth, Maria Wong, D. Monty Wood, Norman Woodley, Tiffany Yau, Thomas J. Zavortink, Manuel A. Zumbado. 2018. Remarkable fly (Diptera) diversity in a patch of Costa Rican cloud forest: Why inventory is a vital science
Brian V. Brown, Art Borkent, Peter H. Adler, Dalton de Souza Amorim, Kevin Barber, Daniel Bickel, Stephanie Boucher, Scott E. Brooks, John Burger, Zelia L. Burington, Renato S. Capellari, Daniel N. R. Costa, Jeffrey M. Cumming, Greg Curler, Carl W. Dick, John H. Epler, Eric Fisher, Stephen D. Gaimari, Jon Gelhaus, David A. Grimaldi, John Hash, Martin Hauser, Heikki Hippa, Sergio Ibanez-Bernal, Mathias Jaschhof, Elena P. Kameneva, Peter H. Kerr, Valery Korneyev, Cheslavo A. Korytkowski, Giar-Ann Kung, Gunnar Mikalsen Kvifte, Owen Lonsdale, Stephen A. Marshall, Wayne Mathis, Verner Michelsen, Stefan Naglis, Allen L. Norrbom, Steven Paiero, Thomas Pape, Alessandre Pereira-Colavite, Marc Pollet, Sabrina Rochefort, Alessandra Rung, Justin B. Runyon, Jade Savage, Vera C. Silva, Bradley J. Sinclair, Jeffrey H. Skevington, John O. Stireman, John Swann, F. Christian Thompson, Pekka Vilkamaa, Terry Wheeler, Terry Whitworth, Maria Wong, D. Monty Wood, Norman Woodley, Tiffany Yau, Thomas J. Zavortink, Manuel A. Zumbado. 2018. Comprehensive inventory of true flies (Diptera) at a tropical site
Mary A. Jamieson, Laura A. Burkle, Jessamyn S. Manson, Justin B. Runyon, Amy M. Trowbridge, Joseph Zientek. 2017. Global change effects on plant-insect interactions: The role of phytochemistry
Laura A. Burkle, Justin B. Runyon. 2017. The smell of environmental change: Using floral scent to explain shifts in pollinator attraction
Justin B. Runyon. 2017. Invasive Species Science Update (No. 9)
Justin B. Runyon. 2016. A dolichopodid hotspot: Montana's Milligan Creek Canyon
Thomas Pape, Allen Allison, Daniel J. Bickel, Torsten Dikow, Thomas Donegan, Donald W. Duszynski, Magdi S. El-Hawagry, Neal L. Evenhuis, Daphne G. Fautin, Stephen D. Gaimari, Babak Gharali, Dale E. Greenwalt, Hinrich Kaiser, Ashley H. Kirk-Spriggs, Gerardo Lamas, Owen Lonsdale, Christopher Mah, Stephen A. Marshall, Rudolf Meier, Michael Ohl, David J. Patterson, Lyubomir Penev, N. Dean Pentcheff, Richard L. Pyle, Daniel J. Rubinoff, Justin B. Runyon, Oliver Tallowin, Stephen Thorpe, Bo Wang Nanjing, Francisco Welter-Schultes, Douglas Yanega, Ding Yang, Gang Yao Hangzhou, Norine W. Yeung. 2016. Species can be named from photos
Tatyana A. Rand, Wendell L. Morrill, Justin B. Runyon, Kim A. Hoelmer, Thomas G. Shanower, Jeffrey L. Littlefield, David K. Weaver. 2016. Assessing phenological synchrony between the Chinese sawfly, Cephus fumipennis, its egg-larval parasitoid, Collyria catoptron, and the North American sawfly, Cephus cinctus: Implications for biological control
A. D. Giunta, Justin B. Runyon, M. J. Jenkins, M. Teich. 2016. Volatile and within-needle terpene changes to Douglas-fir trees associated with Douglas-fir beetle (Coleoptera: Curculionidae) attack
Justin B. Runyon, Jennifer L. Birdsall. 2016. Costs of induced defenses for the invasive plant houndstongue (Cynoglossum officinale L.) and the potential importance for weed biocontrol
Deidre M. Jaeger, Justin B. Runyon, Bryce A. Richardson. 2016. Signals of speciation: Volatile organic compounds resolve closely related sagebrush taxa, suggesting their importance in evolution
Dean E. Pearson, Yvette K. Ortega, Justin B. Runyon, Jack L. Butler. 2016. Secondary invasion: The bane of weed management
Laura A. Burkle, Justin B. Runyon. 2016. Drought and leaf herbivory influence floral volatiles and pollinator attraction
Deborah M. Finch, Rosemary L. Pendleton, Matt C. Reeves, Jeffrey E. Ott, Francis F. Kilkenny, Jack L. Butler, Jacqueline P. Ott, Jeremy R. Pinto, Paulette L. Ford, Justin B. Runyon, Mark A. Rumble, Stanley G. Kitchen. 2016. Rangeland drought: Effects, restoration, and adaptation [Chap. 8]
Christopher J. Fettig, Kenneth E. Gibson, Carl L. Jørgensen, Steven Munson, Jose F. Negrón, Justin B. Runyon, Brytten E. Steed. 2015. Impacts of mountain pine beetle (Dendroctonus ponderosae) outbreaks on forest conditions in the Intermountain West
Kas Dumroese, Tara Luna, Bryce A. Richardson, Francis F. Kilkenny, Justin B. Runyon. 2015. Conserving and restoring habitat for Greater Sage-Grouse and other sagebrush-obligate wildlife: The crucial link of forbs and sagebrush diversity
Curtis A. Gray, Justin B. Runyon, Michael J. Jenkins, Andrew D. Giunta. 2015. Mountain pine beetles use volatile cues to locate host limber pine and avoid non-host Great Basin bristlecone pine
Justin B. Runyon. 2015. Haromyia, a new genus of long-legged flies from Dominica (Diptera: Dolichopodidae)
Justin B. Runyon, Michael A. Ivie, Neal L. Evenhuis. 2015. Liancalus Loew, 1857 and Scellus Loew, 1857 (Insecta, Diptera, DOLICHOPODIDAE): Proposed conservation of the names by designation of Dolichopus regius Fabricius, 1805 as type species for Anoplomerus Rondani, 1856 [Case 3681]
Justin B. Runyon, Richard L. Hurley. 2015. A revision of the Nearctic species of Liancalus (Diptera: Dolichopodidae)
Wesley G. Page, Michael J. Jenkins, Justin B. Runyon. 2014. Spruce beetle-induced changes to Engelmann spruce foliage flammability
Nancy E. Gillette, David L. Wood, Sarah J. Hines, Justin B. Runyon, Jose Negron. 2014. The once and future forest: Consequences of mountain pine beetle treatment decisions
Michael J. Jenkins, Justin B. Runyon, Christopher J. Fettig, Wesley G. Page, Barbara J. Bentz. 2014. Interactions among the mountain pine beetle, fires, and fuels
J. L. Littlefield, G. Markin, J. Kashefi, A. de Meij, Justin B. Runyon. 2013. The release and recovery of Bradyrrhoa gilveolella on rush skeletonweed in southern Idaho
Wesley G. Page, Michael J. Jenkins, Justin B. Runyon. 2012. Mountain pine beetle attack alters the chemistry and flammability of lodgepole pine foliage
Justin B. Runyon, Jack L. Butler, Megan M. Friggens, Susan E. Meyer, Sharlene E. Sing. 2012. Invasive species and climate change (Chapter 7)
Justin B. Runyon. 2012. The Nearctic species of Telmaturgus (Diptera: Dolichopodidae)
Jennifer L. Birdsall, Ward McCaughey, Justin B. Runyon. 2012. Roads impact the distribution of noxious weeds more than restoration treatments in a lodgepole pine forest in Montana, U.S.A
Justin B. Runyon. 2010. A new synonymy in Erebomyia (Diptera: Dolichopodidae)
Justin B. Runyon, Mark C. Mescher, Consuelo M. De Moraes. 2010. Plant defenses against parasitic plants show similarities to those induced by herbivores and pathogens
Justin B. Runyon, Harold Robinson. 2010. Hurleyella, a new genus of Nearctic Dolichopodidae (Diptera)
Justin B. Runyon, Mark C. Mescher, Gary W. Felton, Consuelo M. De Moraes. 2010. Parasitism by Cuscuta pentagona sequentially induces JA and SA defence pathways in tomato
Justin B. Runyon, John F. Tooker, Mark C. Mescher, Consuelo M. De Moraes. 2009. Parasitic plants in agriculture: Chemical ecology of germination and host-plant location as targets for sustainable control: A review
David K. Weaver, Micaela Buteler, Megan L. Hofland, Justin B. Runyon, Christian Nansen, Luther E. Talbert, Peggy Lamb, Gregg R. Carlson. 2009. Cultivar preferences of ovipositing wheat stem sawflies as influenced by the amount of volatile attractant
Richard L. Hurley, Justin B. Runyon. 2009. A review of Erebomyia (Diptera: Dolichopodidae), with descriptions of three new species
Justin B. Runyon. 2008. New species of long-legged flies (Diptera: Dolichopodidae) from central Pennsylvania
Justin B. Runyon. 2008. The cingulum: A unique structure of some Dolichopodidae
Justin B. Runyon. 2008. Richard L. Hurley, 1934-2008
Justin B. Runyon, Mark C. Mescher, Consuelo M. De Moraes. 2008. Parasitism by Cuscuta pentagona attenuates host plant defenses against insect herbivores
Mark C. Mescher, Justin B. Runyon, Consuelo M. De Moraes. 2006. Plant host finding by parasitic plants: A new perspective on plant to plant communication
Justin B. Runyon, Mark C. Mescher, Consuelo M. De Moraes. 2006. Volatile chemical cues guide host location and host selection by parasitic plants
Christian Nansen, David K. Weaver, Sharlene E. Sing, Justin B. Runyon, Wendell L. Morrill, Matthew J. Grieshop, Courtney L. Shannon, Megan L. Johnson. 2005. Within-field spatial distribution of Cephus cinctus (Hymenoptera: Cephidae) larvae in Montana wheat fields
David K. Weaver, Christian Nansen, Justin B. Runyon, Sharlene E. Sing, Wendell L. Morrill. 2005. Spatial distributions of Cephus cinctus Norton (Hymenoptera: Cephidae) and its braconid parasitoids in Montana wheat fields
Justin B. Runyon, Richard L. Hurley. 2004. A new genus of long-legged flies displaying remarkable wing directional asymmetry
Justin B. Runyon, Richard L. Hurley. 2003. Revision of the Nearctic species of Nepalomyia Hollis (= Neurigonella Robinson) (Diptera: Dolichopodidae: Peloropeodinae) with a world catalog
Bryn H. Daisy, Gary A. Strobel, Uvidelio Castillo, David Ezra, Joe Sears, David K. Weaver, Justin B. Runyon. 2002. Naphthalene, an insect repellent, is produced by Muscodor vitigenus, a novel endophytic fungus

University of Virginia's College at Wise, VA, B.S., Biology and Mathematics, 1998
Montana State University, Bozeman, MT, M.S., Entomology, 2001
Pennsylvania State University, University Park, PA, Ph.D., Entomology, 2008

Last updated March 10, 2026

Justin B. Runyon, PhD

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