Kristine L. Pilgrim

Laboratory Supervisor, Biological Scientist

Rocky Mountain Research Station

0000-0003-3726-6980

kristine.pilgrim@usda.gov

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Education

Treesearch Publications

Kalon C. Baughan, Bret H. Davis, Kristine L. Pilgrim, Anne A. Carlson, Sergio E. Morales, Phillip W. Ramsey, Michael K. Schwartz, Mikaela G. Howie. 2025. A portable structure for identifying wolverines and Canada lynx using integrated cameras and hair snags
Alexandra Fraik, Kristine L. Pilgrim, Cory E. Mosby, Richard Weir, Cameron L. Heusser, Nathan Kluge, Michael K. Schwartz. 2025. The genetic implications of translocations on Fisher (Pekania pennanti) populations in Montana and Idaho
Imogene A. Cancellare, Byron Weckworth, Anthony Caragiulo, Kristine L. Pilgrim, Thomas M. McCarthy, Alisher Abdullaev, George Amato, Xiaoxing Bian, Elena Bykova, Isabela Dias-Freedman, Mariya Gritsina, Lauren M. Hennelly, Safia Janjua, Orjan Johansson, Shannon Kachel, Alexander Karnaukhov, Miroslav Korablev, Zairbek Kubanychbekov, Rahim Kulenbekov, Xuchang Liang, Purevjav Lkhagvajav, Tara K. Meyer, Bariushaa Munkhtsog, Bayaraa Munkhtsog, Muhammad A. Nawaz, Stephane Ostrowski, Mikhail Paltsyn, Andrey Poyarkov, Salisa Rabinowitz, Thomas Rooney, Tatjana Rosen, Viatcheslav V Rozhnov, Benjamin N. Sacks, Michael K. Schwartz, Kyle P. McCarthy. 2024. Snow leopard phylogeography and population structure supports two global populations with single refugial origin
L. Mark Elbroch, Byron V Weckworth, Kristine L. Pilgrim, Omar Ohrens, Nicolas Lagos, Stephanny Arroyo-Arce, Mauricio Montt, Dania Goic, Michael K. Schwartz. 2024. An initial genetic assessment of the emblematic pumas of the Torres del Paine UNESCO Biosphere Reserve
Casey C. Day, Erin L. Landguth, Michael A. Sawaya, Anthony P. Clevenger, Robert A. Long, Zachary Holden, Jocelyn R. Akins, Robert B. Anderson, Keith B. Aubry, Mirjam Barrueto, Nichole L. Bjornlie, Jeffrey P. Copeland, Jason T. Fisher, Anne Forshner, Justin A. Gude, Doris Hausleitner, Nichole A. Heim, Kimberly S. Heinemeyer, Anne Hubbs, Robert M. Inman, Scott Jackson, Michael Jokinen, Nathan P. Kluge, Andrea Kortello, Deborah L. Lacroix, Luke Lamar, Lisa I Larson, Jeffrey C. Lewis, Dave Lockman, Michael K. Lucid, Paula MacKay, Audrey J. Magoun, Michelle L. McLellan, Katie Moriarty, Cory E. Mosby, Garth Mowat, Clifford G. Nietvelt, David Paetkau, Eric C. Palm, Kylie J. S. Paul, Kristine L. Pilgrim, Catherine M. Raley, Michael K. Schwartz, Matthew A. Scrafford, John R. Squires, Zachary J. Walker, John S. Waller, Richard D. Weir, Katherine Zeller. 2024. Genetic connectivity of wolverines in western North America
Dave P. Onorato, Mark W. Cunningham, Mark Lotz, Marc Criffield, David Shindle, Annette Johnson, Bambi C. F. Clemons, Colin P. Shea, Melody E. Roelke‑Parker, Warren E. Johnson, Brett T. McClintock, Kristine L. Pilgrim, Michael K. Schwartz, Madan K. Oli. 2024. Multi‑generational benefits of genetic rescue
Helen E. Chmura, Lucretia E. E. Olson, Remi Murdoch, Alexandra Fraik, Scott Jackson, Kevin S. McKelvey, Rex Koenig, Kristine L. Pilgrim, Nicholas DeCesare, Michael K. Schwartz. 2024. Climate change differentially alters distribution of two marten species in a hybrid zone
Richard D. Weir, Andrew M. Rankin, Lacy Robinson, Kristine L. Pilgrim, Michael K. Schwartz, Michael K. Lucid. 2024. Genetic structuring of fishers in British Columbia, Canada: Implications for population conservation and management
Daniel Vanderpool, Taylor Wilcox, Michael K. Young, Kristine L. Pilgrim, Michael K. Schwartz. 2024. Simultaneous species detection and discovery with environmental DNA metabarcoding: A freshwater mollusk case study
Gretchen H. Roffler, Kristine L. Pilgrim, Benjamin C. Williams. 2024. Patterns of wolf dispersal respond to harvest density across an island complex
Melissa Reynolds-Hogland, Carly Brooks, Alan B. Ramsey, John S. Hogland, Kristine L. Pilgrim, Cory L. Engkjer, Philip W. Ramsey. 2024. Long-term video and genetic data yield insights into complex sociality of a solitary large carnivore
L. M. Elbroch, S. H. Williams, O. Ohrens, Kristine L. Pilgrim, A. Moeller, S. Arroyo-Arce, M. Parker, D. Goic, H. Robinson, Michael K. Schwartz. 2024. Comparing abundance estimates of a cryptic carnivore in southern Patagonia using two experimental methods
Gretchen H. Roffler, Kristine L. Pilgrim, Katherine Zarn, Michael K. Schwartz, Taal Levi. 2023. Variation in adult and pup wolf diets at natal den sites is influenced by forest composition and configuration
Francesca Marucco, Maria V Boiani, Pierre Dupont, Cyril Milleret, Elisa Avanzinelli, Kristine L. Pilgrim, Michael K. Schwartz, Achaz von Hardenberg, Domenica Serena Perrone, Olivier P. Friard, Arianna Menzano, Francesco Bisi, Umberto Fattori, Michela Tomasella, Sonia Calderola, Sabrina Carolfi, Piero Ferrari, Christian Chioso, Fabrizio Truc, Giulia Bombieri, Luca Pedrotti, Davide Righetti, Pierluigi L. Acutis, Fabio Guglielmo, Heidi C. Hauffe, Chiara Rossi, Romolo Caniglia, Paola Aragno, Valentina La Morgia, Piero Genovesi, Richard Bischof. 2023. A multidisciplinary approach to estimating wolf population size for long-term conservation
Kristine L. Pilgrim, Rebecca E. Green, Kathryn L. Purcell, Taylor Wilcox, Eric L. McGregor, Larissa E. Gleason, Samuel K. Wasser, Michael K. Schwartz. 2023. Shifts in fisher (Pekania pennanti) diet in response to climate-induced tree mortality in California assessed with DNA metabarcoding
Melissa Reynolds-Hogland, Alan B. Ramsey, Carly Muench, Kirsten Terkildsen, Kristine L. Pilgrim, Cory L. Engkjer, Philip W. Ramsey. 2023. Video-documentation of true and borderline tool use by wild American black bears
Michael K. Young, Rebecca Smith, Kristine L. Pilgrim, Daniel J. Isaak, Kevin S. McKelvey, Sharon L. Payne, Jacob Egge, Michael K. Schwartz. 2022. A molecular taxonomy of Cottus in western North America
Melissa Reynolds-Hogland, Alan B. Ramsey, Carly Muench, Kristine L. Pilgrim, Cory L. Engkjer, Grace Erba, Philip W. Ramsey. 2022. Integrating video and genetic data to estimate annual age-structured apparent survival of American black bears
Francesca Marucco, Kristine L. Pilgrim, Elisa Avanzinelli, Michael K. Schwartz, Luca Rossi. 2022. Wolf dispersal patterns in the Italian Alps and implications for wildlife diseases spreading
Jessie Golding, Cory R. Davis, Luke Lamar, Scott Tomson, Carly Lewis, Kristine L. Pilgrim, Mark Ruby, Mike Mayernik, Kevin S. McKelvey. 2022. Targeted efforts are more effective than combined approaches for sampling two rare carnivores
Melissa J. Reynolds-Hogland, Alan B. Ramsey, Carly Muench, Kristine L. Pilgrim, Cory L. Engkjer, Philip W. Ramsey. 2022. Age-specific, population-level pedigree of wild black bears provides insights into reproduction, paternity, and maternal effects on offspring apparent survival
David S. Green, Marie E. Martin, Roger A. Powell, Eric L. McGregor, Mourad Gabriel, Kristine L. Pilgrim, Michael K. Schwartz, Sean M. Matthews. 2022. Mixed-severity wildfire and salvage logging affect the populations of a forest-dependent carnivoran and a competitor
Michael K. Young, Rebecca Smith, Kristine L. Pilgrim, Michael K. Schwartz. 2021. Molecular species delimitation refines the taxonomy of native and nonnative physinine snails in North America
Melissa J. Reynolds-Hogland, Alan B. Ramsey, August T. Seward, Kristine L. Pilgrim, Cory L. Engkjer, Philip W. Ramsey. 2020. Response of a remnant marmot population to habitat enhancement yields insights into marmot ecology
Shannon M. Barber-Meyer, Joseph Dysthe, Kristine L. Pilgrim. 2020. Testing environmental DNA from wolf snow tracks for species, sex, and individual identification
Kevin S. McKelvey, Corey Kallstrom, Jeri Ledbetter, Donald W. Sada, Kristine L. Pilgrim, Michael K. Schwartz. 2020. An inventory of springsnails (Pyrgulopsis spp.) in and adjacent to the Spring Mountains, Nevada
Michael K. Schwartz, Ashley Walters, Kristine L. Pilgrim, Katie Moriarty, K. M. Slauson, Bill Zielinski, Keith B. Aubry, B. N. Sacks, Katherine Zarn, C. B. Quinn, Michael K. Young. 2020. Pliocene-early Pleistocene geological events structure Pacific martens (Martes caurina)
Nicholas J. DeCesare, Byron V Weckworth, Kristine L. Pilgrim, Andrew B. D. Walker, Eric J. Bergman, Kassidy E. Colson, Rob Corrigan, Richard B. Harris, Mark Hebblewhite, Brett R. Jesmer, Jesse R. Newby, Jason R. Smith, Rob B. Tether, Timothy P. Thomas, Michael K. Schwartz. 2020. Phylogeography of moose in western North America
Patricia J. Happe, Kurt J. Jenkins, Rebecca M. Mccaffery, Jeffrey C. Lewis, Kristine L. Pilgrim, Michael K. Schwartz. 2020. Occupancy patterns in a reintroduced fisher population during reestablishment
Daniel B. Thompson, Kevin S. McKelvey, Paul van Els, Gretchen Andrew, Paula Jacoby‑Garrett, Matt Glenn, Corey Kallstrom, Kristine L. Pilgrim, Paul A. Opler. 2020. Conserve the eco-evolutionary dynamic, not the subspecies: Phenological divergence and gene flow between temporal cohorts of Euphilotes ancilla endemic to southern Nevada
Laurel E. Peelle, Aaron J. Wirsing, Kristine L. Pilgrim, Michael K. Schwartz. 2019. Identifying predators from saliva at kill sites with limited remains
Sean M. Johnson-Bice, Steve K. Windels, Kristine L. Pilgrim. 2019. Observation and DNA confirmation of a fisher (Pekania pennanti) hunting and killing a muskrat (Ondatra zibethicus)
Katie Moriarty, Keith B. Aubry, Connor N. Morozumi, Betsy L. Howell, Patricia J. Happe, Kurt J. Jenkins, Kristine L. Pilgrim, Michael K. Schwartz. 2019. Status of Pacific Martens (Martes caurina) on the Olympic Peninsula, Washington
Gretchen H. Roffler, Jason N. Waite, Kristine L. Pilgrim, Katherine E. Zarn, Michael K. Schwartz. 2019. Estimating abundance of a cryptic social carnivore using spatially explicit capture-recapture
Michael K. Young, Rebecca J. Smith, Kristine L. Pilgrim, Matthew P. Fairchild, Michael K. Schwartz. 2019. Integrative taxonomy refutes a species hypothesis: The asymmetric hybrid origin of Arsapnia arapahoe (Plecoptera, Capniidae)
Kevin S. McKelvey, Jessie Golding, Daniel H. Mason, Joseph Dysthe, Kristine L. Pilgrim, John R. Squires, Thomas W. Franklin, Keith B. Aubry, Robert A. Long, Samuel Greaves, Catherine M. Raley, Scott Jackson, Paula MacKay, Joshua Lisbon, Joel D. Sauder, Michael T. Pruss, Don Heffington, Michael K. Schwartz. 2019. Using environmental DNA methods to improve winter surveys for rare carnivores: DNA from snow and improved noninvasive techniques
Michael K. Young, Kevin S. McKelvey, Tara Jennings, Katie Carter, Richard Cronn, Ernest R. Keeley, Janet L. Loxterman, Kristine L. Pilgrim, Michael K. Schwartz. 2018. The phylogeography of westslope cutthroat trout
Christopher P. Kirol, Kristine L. Pilgrim, Andrew L. Sutphin, Thomas L. Maechtle. 2018. Using DNA from hairs left at depredated greater sage-grouse nests to detect mammalian nest predators
Rebeca L. Juarez, Michael K. Schwartz, Kristine L. Pilgrim, Daniel J. Thompson, Stephanie A. Tucker, Joshua Smith, Jonathan A. Jenks. 2016. Assessing temporal genetic variation in a cougar population: Influence of harvest and neighboring populations
Heidi Kretser, Michale Glennon, Alice Whitelaw, Aimee Hurt, Kristine L. Pilgrim, Michael K. Schwartz. 2016. Scat-detection dogs survey low density moose in New York
Michael K. Young, Daniel J. Isaak, Kevin S. McKelvey, Taylor M. Wilcox, Daniel M. Bingham, Kristine L. Pilgrim, Kellie Carim, Matthew R. Campbell, Matthew P. Corsi, Dona Horan, David E. Nagel, Michael K. Schwartz. 2016. Climate, demography, and zoogeography predict introgression thresholds in salmonid hybrid zones in Rocky Mountain streams
Gretchen H. Roffler, Michael K. Schwartz, Kristine L. Pilgrim, Sandra L. Talbot, George K. Sage, Layne G. Adams, Gordon Luikart. 2016. Identification of landscape features influencing gene flow: How useful are habitat selection models?
Jason E. Hawley, Paul W. Rego, Adrian P. Wydeven, Michael K. Schwartz, Tabitha C. Viner, Roland Kays, Kristine L. Pilgrim, Jonathan A. Jenks. 2016. Long-distance dispersal of a subadult male cougar from South Dakota to Connecticut documented with DNA evidence
Kevin S. McKelvey, Michael K. Young, Taylor M. Wilcox, Daniel M. Bingham, Kristine L. Pilgrim, Michael K. Schwartz. 2016. Patterns of hybridization among cutthroat trout and rainbow trout in northern Rocky Mountain streams
K. M. Proffitt, Joshua F. Goldberg, M. Hebblewhite, R. Russell, B. S. Jimenez, H. S. Robinson, Kristine L. Pilgrim, Michael K. Schwartz. 2015. Integrating resource selection into spatial capture-recapture models for large carnivores
Gretchen H. Roffler, Sandra L. Talbot, Gordon Luikart, George K. Sage, Kristine L. Pilgrim, Layne G. Adams, Michael K. Schwartz. 2014. Lack of sex-biased dispersal promotes fine-scale genetic structure in alpine ungulates
Kevin S. McKelvey, Keith B. Aubry, Neil J. Anderson, Anthony P. Clevenger, Jeffrey P. Copeland, Kimberley S. Heinemeyer, Robert M. Inman, John R. Squires, John S. Waller, Kristine L. Pilgrim, Michael K. Schwartz. 2014. Recovery of wolverines in the western United States: Recent extirpation and recolonization or range retraction and expansion?
Michael Lemoine, Michael K. Young, Kevin S. McKelvey, Lisa Eby, Kristine L. Pilgrim, Michael K. Schwartz. 2014. Cottus schitsuumsh, a new species of sculpin (Scorpaeniformes: Cottidae) in the Columbia River basin, Idaho-Montana, USA
Michael K. Schwartz, Joel Saunder, Kristine L. Pilgrim, Ray Vinkey, Michael K. Lucid, Sean A. Parks, Nathan Albrecht. 2013. Fisher population and landscape genetics
Kevin S. McKelvey, Jennifer E. Ramirez, Kristine L. Pilgrim, Samuel A. Cushman, Michael K. Schwartz. 2013. Genetic sampling of Palmer's chipmunks in the Spring Mountains, Nevada
Michael Lucid, L. Robinson, Samuel A. Cushman, L. Allen, Michael K. Schwartz, Kristine L. Pilgrim. 2013. Status of fisher in the northern Idaho panhandle and adjacent mountain ranges
Bill Zielinski, Ric Schlexer, T. Luke George, Kristine L. Pilgrim, Michael K. Schwartz. 2013. Estimating abundance and survival in the endangered Point Arena Mountain beaver using noninvasive genetic methods
Michael K. Young, Kevin S. McKelvey, Kristine L. Pilgrim, Michael K. Schwartz. 2013. DNA barcoding at riverscape scales: Assessing biodiversity among fishes of the genus Cottus (Teleostei) in northern Rocky Mountain streams
Jody M. Tucker, Michael K. Schwartz, Richard L. Truex, Kristine L. Pilgrim, Fred W. Allendorf. 2012. Historical and contemporary DNA indicate fisher decline and isolation occurred prior to the European settlement of California
Kristine L. Pilgrim, Bill Zielinski, Ric Schlexer, Michael K. Schwartz. 2012. Development of a reliable method for determining sex for a primitive rodent, the Point Arena mountain beaver (Aplodontia rufa nigra)
Bill Zielinski, Ric Schlexer, Sean A. Parks, Kristine L. Pilgrim, Michael K. Schwartz. 2012. Small geographic range but not panmictic: how forests structure the endangered Point Arena mountain beaver (Aplodontia rufa nigra)
Audrey J. Magoun, Clinton D. Long, Michael K. Schwartz, Kristine L. Pilgrim, Richard E. Lowell, Patrick Valkenburg. 2011. Integrating motion-detection cameras and hair snags for wolverine identification
Brian Knaus, Richard Cronn, Aaron Liston, Kristine L. Pilgrim, Michael K. Schwartz. 2011. Mitochondrial genome sequences illuminate maternal lineages of conservation concern in a rare carnivore
Todd J. Brinkman, David K. Person, Michael K. Schwartz, Kristine L. Pilgrim, Kevin E. Colson, Kris J. Hundertmark. 2010. Individual identification of Sitka black-tailed deer (Odocoileus hemionus sitkensis) using DNA from fecal pellets
Todd J. Brinkman, Michael K. Schwartz, David K. Person, Kristine L. Pilgrim, Kris J. Hundertmark. 2009. Effects of time and rainfall on PCR success using DNA extracted from deer fecal pellets
Katie M. Moriarty, Bill Zielinski, Armand G Gonzales, Todd E. Dawson, Kristie M. Boatner, Craig A Wilson, Ric Schlexer, Kristine L. Pilgrim, Jeff Copeland, Michael K. Schwartz. 2009. Wolverine confirmation in California after nearly a century: native or long-distance immigrant
Jessica A. Homyack, Jennifer H. Vashon, Cade Libby, Edward L. Lindquist, Steve Loch, Donald F. McAlpine, Kristine L. Pilgrim, Michael K. Schwartz. 2008. Canada lynx-bobcat (Lynx canadensis x L. rufus) hybrids at the southern periphery of lynx range in Maine, Minnesota and New Brunswick
Michael K. Schwartz, Kristine L. Pilgrim, Kevin S. McKelvey, Pilar T. Rivera, Leonard F. Ruggiero. 2007. DNA markers for identifying individual snowshoe hares using field-collected pellets
Michael K. Schwartz, Keith B. Aubry, Kevin S. McKelvey, Kristine L. Pilgrim, Jeffrey P. Copeland, John R. Squires, Robert M. Inman, Samantha M. Wisely, Leonard F. Ruggiero. 2007. Inferring geographic isolation of wolverines in California using historical DNA
Bill Zielinski, Fredrick V. Schlexer, Kristine L. Pilgrim, Michael K. Schwartz. 2006. The efficacy of wire and glue hair snares in identifying mesocarnivores
Kristine L. Pilgrim, Bill Zielinski, Mary J. Mazurek, Ric Schlexer, Michael K. Schwartz. 2006. Development and characterization of microsatellite markers in the Point Arena mountain beaver Aplodontia rufa nigra
Ray S. Vinkey, Michael K. Schwartz, Kevin S. McKelvey, Kerry R. Foresman, Kristine L. Pilgrim, Brian J. Giddings, Eric C. Lofroth. 2006. When reintroductions are augmentations: the genetic legacy of the fisher (Martes pennanti) in Montana