Melody Keena, PhD

Research Entomologist

Northern Research Station

0000-0003-3099-6243

melody.keena@usda.gov

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Treesearch Publications

Melody Keena, R. Talbot Trotter, III, Amanda D. Roe, David R. Coyle. 2026. New Insights on the Ecology, Economics, and Management of Anoplophora Longhorned Beetles (Coleoptera: Cerambycidae: Lamiinae)
Melody Keena, Nooshin Zandi-Sohani, Jessica Y. Richards, Juan Shi, Yuri N Baranchikov. 2026. Flighted spongy moths (Lepidoptera: Erebidae), defoliating invasive forest pests
Sabrina Osowiecki, Melody Keena. 2025. Effects of dietary moisture content and cut wood of different species on survival and weight of Asian longhorned beetle larvae (Coleoptera: Cerambycidae)
Nooshin Zandi-Sohani, Melody Keena, Michael R. Gallagher, Anthony Cullen, Dong H. Cha. 2025. Heat treatments to kill eggs of two invasive forest insects: Lycorma delicatula (Hemiptera: Fulgoridae) and Lymantria dispar (Lepidoptera: Erebidae)
Mingming Cui, Amanda D. Roe, Brian Boyle, Melody Keena, Yunke Wu, W. Evan Braswell, Michael T. Smith, Ben Gasman, Juan Shi, Marion Javal, Geraldine Roux, Jean J. Turgeon, Richard Hamelin, Ilga Porth. 2024. Tracking the North American Asian Longhorned Beetle Invasion with Genomics
Melody Keena. 2024. Effects of temperature on the survival of spotted lanternfly active life stages when held without food
Gwylim S. Blackburn, Christopher I Keeling, Julien Prunier, Melody Keena, Catherine Béliveau, Richard Hamelin, Nathan P. P. Havill, Francois Olivier Hebert, Roger C. Levesque, Michel Cusson, Ilga Porth. 2024. Genetics of flight in spongy moths (Lymantria dispar ssp.): functionally integrated profiling of a complex invasive trait
Kishan R. Sambaraju, Vivek Srivastava, Brittany S. Barker, Melody Keena, Michael D. Ormsby, Allan L. Carroll. 2024. Editorial: Forest insect invasions – risk mapping approaches and applications
Albert (Bud) E. Mayfield, Tonya D. Bittner, Nicholas J. Dietschler, Joseph S. Elkinton, Nathan P. P. Havill, Melody Keena, David L. Mausel, James R. Rhea, Scott M. Salom, Mark C. Whitmore. 2023. Biological control of hemlock woolly adelgid in North America: History, status, and outlook.
Haiwen Qin, Huachao Xu, Arnaud Capron, Ilga Porth, Mingming Cui, Melody Keena, Xiaofang Deng, Juan Shi, Richard C. Hamelin. 2023. Is there hybridization between 2 species of the same genus in sympatry?—The genetic relationships between Anoplophora glabripennis, Anoplophora chinensis, and putative hybrids
Devin Kreitman, Melody Keena, Anne L. Nielsen, George Hamilton. 2023. The impact of host plant species on instar duration and body weight of nymphal Lycorma delicatula
Melody Keena, George Hamilton, Devin Kreitman. 2023. The potential climatic range of spotted lanternfly may be broader than previously predicted
Devin Kreitman, Melody Keena, Anne Nielsen, George Hamilton. 2022. The effect of host plant on the nymphal development of spotted lanternfly
Samita Limbu, Melody Keena, Nicholas Dietschler, Katharine O'Connor, Mark C. Whitmore. 2022. Estivation and Postestivation Development of Hemlock Woolly Adelgid (Adelges tsugae) (Hemiptera: Adelgidae) at Different Temperatures
Laura J. Nixon, Sharon Jones, Andrew C. Dechaine, Dalton Ludwick, Mauri Hickin, Liam Sullivan, Johanna E. Elsensohn, Juli Gould, Melody Keena, Thomas Kuhar, Douglas G. Pfeiffer, Tracy C. Leskey. 2022. Development of rearing methodology for the invasive Spotted Lanternfly, Lycorma delicatula (Hemiptera: Fulgoridae)
Melody Keena. 2022. Effects of Dietary Iron Quantities and Sources on Lymantria dispar (Lepidoptera: Erebidae) Survival and Development
Mingming Cui, Yunke Wu, Marion Javal, Isabelle Giguère, Géraldine Roux, Jose A. Andres, Melody Keena, Juan Shi, Baode Wang, Evan Braswell, Scott E. Pfister, Richard Hamelin, Amanda Roe, Ilga Porth. 2022. Genome‐scale phylogeography resolves the native population structure of the Asian longhorned beetle, Anoplophora glabripennis (Motschulsky)
Melody Keena, Jessica Y. Richards. 2021. Effects of Temperature on Anoplophora chinensis (Coleoptera: Cerambycidae) Larvae and Pupae
Xingeng Wang, Melody Keena. 2021. Hybridization Potential of Two Invasive Asian Longhorn Beetles
Melody Keena, Paul Moore, Gregg Bradford. 2021. Effects of Temperature on Anoplophora chinensis (Coleoptera: Cerambycidae) Adult Survival, Reproduction, and Egg Hatch
Vivek Srivastava, Melody Keena, Galen E. Maennicke, Richard C. Hamelin, Verena C. Griess. 2021. Potential Differences and Methods of Determining Gypsy Moth Female Flight Capabilities: Implications for the Establishment and Spread in Novel Habitats
Melody Keena, Anne L. Nielsen. 2021. Comparison of the Hatch of Newly Laid Lycorma delicatula (Hemiptera: Fulgoridae) Eggs From the United States After Exposure to Different Temperatures and Durations of Low Temperature
Vivek Srivastava, Amanda D. Roe, Melody Keena, Richard C. Hamelin, Verena C. Griess. 2021. Oh the places they’ll go: improving species distribution modelling for invasive forest pests in an uncertain world
Devin Kreitman, Melody Keena, Anne Nielsen, George Hamilton. 2020. Effects of Temperature on Development and Survival of Nymphal Lycorma delicatula (Hemiptera: Fulgoridae)
Robert L. Harrison, Daniel L. Rowley, Melody Keena. 2020. Pathology and genome sequence of a Lymantria dispar multiple nucleopolyhedrovirus (LdMNPV) isolate from Heilongjiang, China
Vivek Srivastava, Wanwan Liang, Melody Keena, Amanda D. Roe, Richard C. Hamelin, Verena C. Griess. 2020. Assessing Niche Shifts and Conservatism by Comparing the Native and Post-Invasion Niches of Major Forest Invasive Species
Melody Keena, Jessica Y. Richards. 2020. Comparison of Survival and Development of Gypsy Moth Lymantria dispar L. (Lepidoptera: Erebidae) Populations from Different Geographic Areas on North American Conifers
Gwylim Blackburn, Pierre Bilodeau, Tracey Cooke, Mingming Cui, Michel Cusson, Richard Hamelin, Melody Keena, Sandrine Picq, Amanda Roe, Juan Shi, Yunke Wu, Ilga Porth. 2020. An Applied Empirical Framework for Invasion Science: Confronting Biological Invasion Through Collaborative Research Aimed at Tool Production
R. Talbot Trotter, III, Samita Limbu, Kelli Hoover, Hannah Nadel, Melody Keena. 2020. Comparing Asian Gypsy Moth [Lymantria dispar asiatica (Lepidoptera: Erebidae) and L. dispar japonica] Trap Data From East Asian Ports With Lab Parameterized Phenology Models: New Tools and Questions
Vivek Srivastava, Verena C. Griess, Melody Keena. 2020. Assessing the Potential Distribution of Asian Gypsy Moth in Canada: A Comparison of Two Methodological Approaches
Claire Rutledge, Melody Keena. 2019. Mating Behavior and Reproductive Biology of Emerald Ash Borer (Coleoptera: Buprestidae) and Two of Its Native Congeners, the Twolined Chestnut Borer and the Bronze Birch Borer
David Gray, Melody Keena. 2019. A Phenology Model for Asian Gypsy Moth Egg Hatch
Melody Keena, Juan Shi. 2019. Effects of Temperature on First Instar Lymantria (Lepidoptera: Erebidae) Survival and Development With and Without Food
Marion Javal, Alain Roques, Julien Haran, Franck Hérard, Melody Keena, Géraldine Roux. 2019. Complex invasion history of the Asian long-horned beetle: fifteen years after first detection in Europe
S. Limbu, Melody Keena, M.C. Whitmore. 2018. Hemlock Woolly Adelgid (Hemiptera: Adelgidae): A Non-Native Pest of Hemlocks in Eastern North America
Melody Keena, V. Sánchez. 2018. Inter- and Intrasexual Interactions in Anoplophora glabripennis (Coleoptera: Cerambycidae) and the Impact of Different Sex Ratios
Melody Keena. 2018. Factors That Influence Flight Propensity in Anoplophora glabripennis (Coleoptera: Cerambycidae)
Melody Keena, Vicente Sanchez. 2018. Reproductive Behaviors of Anoplophora glabripennis (Coleoptera: Cerambycidae) in the Laboratory
Sandrine Picq, Melody Keena, Nathan P. P. Havill, Don Stewart, Esther Pouliot, Brian Boyle, Roger C. Levesque, Richard C. Hamelin, Michel Cusson. 2018. Assessing the potential of genotyping-by-sequencing-derived single nucleotide polymorphisms to identify the geographic origins of intercepted gypsy moth (Lymantria dispar) specimens: A proof-of-concept study
Samita Limbu, Melody Keena, Fang Chen, Gericke Cook, Hannah Nadel, Kelli Hoover. 2017. Effects of temperature on development of Lymantria dispar asiatica and Lymantria dispar japonica (Lepidoptera: Erebidae)
Robert A. A. Haack, Melody Keena, Dominic Eyre. 2017. Life history and population dynamics of Cerambycidae
Melody Keena. 2017. Laboratory rearing and handling of cerambycids
Alexander P. Kappel, R. Talbot Trotter, III, Melody Keena, John Rogan, Christopher A. Williams. 2017. Mapping of the Asian longhorned beetle's time to maturity and risk to invasion at contiguous United States extent
R. Talbot Trotter, III, Melody Keena. 2016. A variable-instar climate-driven individual beetle-based phenology model for the invasive Asian longhorned beetle (Coleoptera: Cerambycidae)
Fern Graves, Thomas C. Baker, Aijun Zhang, Melody Keena, Kelli Hoover. 2016. Sensory aspects of trail-following behaviors in the Asian longhorned beetle, Anoplophora glabripennis
Harrison Robert L., Daniel L. Rowley, Melody Keena. 2016. Geographic isolates of Lymantria dispar multiple nucleopolyhedrovirus: Genome sequence analysis and pathogenicity against European and Asian gypsy moth strains
Fang Chen, Juan Shi, Melody Keena. 2016. Evaluation of the effects of light entensity and time interval after the start of scotophase on the female flight propensity of Asian gypsy moth (Lepidoptera: Erebidae)
Daniela Pilarska, Ann E. Hajek, Melody Keena, Andreas Linde, Manana Kereselidze, Georgi Georgiev, Margarita Georgieva, Plamen Mirchev, Danail Takov, Slavimira Draganova. 2016. Susceptibility of larvae of nun moth, Lymantria monacha (Linnaeus 1758) (Lepidoptera), to the entomopathogenic fungus, Entomophaga maimaiga Humber, Shimazu and Soper (Entomophthorales) under laboratory and field conditions
Fang Chen, Youqing Luo, Melody Keena, Ying Wu, Peng Wu, Juan Shi. 2015. DNA barcoding of gypsy moths from China (Lepidoptera: Erebidae) reveals new haplotypes and divergence patterns within gypsy moth subspecies
Samita Limbu, Katie Cassidy, Melody Keena, Patrick Tobin, Kelli Hoover. 2015. Host range specificity of Scymnus camptodromus (Coleoptera: Coccinellidae), a predator of hemlock woolly adelgid (Hemiptera: Adelgidae)
Juan Shi, Fang Chen, Melody Keena. 2015. Differences in wing morphometrics of Lymantria dispar (Lepidoptera: Erebidae) between populations that vary in female flight capability
P.S. Meng, K. Hoover, Melody Keena. 2015. Asian longhorned beetle (Coleoptera: Cerambycidae), an introduced pest of maple and other hardwood trees in North America and Europe
Melody Keena, Hannah Nadel, Juli Gould. 2015. Survival and phenology of Agrilus planipennis (Coleoptera: Buprestidae) reared on a newly developed artificial diet free of host material
Samita Limbu, Melody Keena, David Long, Nancy Ostiguy, Kelli Hoover. 2015. Scymnus camptodromus (Coleoptera: Coccinellidae) larval development and predation of hemlock woolly adelgid (Hemiptera: Adelgidae)
P.S. Meng, R. Talbot Trotter, III, Melody Keena, T.C. Baker, S. Yan, E.G. Schwartzberg, K. Hoover. 2014. Effects of pheromone and plant volatile release rates and ratios on trapping Anoplophora glabripennis (Coleoptera: Cerambycidae) in China
M.E. Nehme, R. Talbot Trotter, III, Melody Keena, C. McFarland, J. Coop, H.M. Hull-Sanders, P. Meng, C.M. De Moraes, M.C. Mescher, K. Hoover. 2014. Development and evaluation of a trapping system for Anoplophora glabripennis (Coleoptera: Cerambycidae) in the United States
Kelli Hoover, Melody Keena, Maya Nehme, Shifa Wang, Peter Meng, Aijun Zhang. 2014. Sex-specific trail pheromone mediates complex mate finding behavior in Anoplophora glabripennis
Robert L. Harrison, Melody Keena, Daniel L. Rowley. 2014. Classification, genetic variation and pathogenicity of Lymantria dispar nucleopolyhedrovirus isolates from Asia, Europe, and North America
Vicente Sanchez, Melody Keena. 2013. Development of the teneral adult Anoplophora glabripennis (Coleoptera: Cerambycidae): time to initiate and completely bore out of maple wood
Melody Keena, R. Talbot Trotter, III, Carole Cheah, Michael Montgomery. 2012. Effects of temperature and photoperiod on the aestivo-hibernal egg diapause of Scymnus camptodromus (Coleoptera: Coccinellidae)
Claire E. Rutledge, Melody Keena. 2012. Mating frequency and fecundity in Agrilus anxius (Coleoptera: Buprestidae)
Claire E. Rutledge, Melody Keena. 2012. Mating frequency and fecundity in the emerald ash borer Agrilus planipennis (Coleoptera: Buprestidae)
Michael Montgomery, Melody Keena. 2011. Scymnus (Neopullus) lady beetles from China
Nathan P. P. Havill, Michael Montgomery, Melody Keena. 2011. Hemlock woolly adelgid and its hemlock hosts: A global perspective
Alexandra N. Whitney, Melody Keena. 2011. Effects of host wood moisture on the life cycle development of the Asian longhorned beetle
Maya Nehme, Melody Keena, Aijun Zhang, Alan Sawyer, Kelli Hoover. 2011. Monitoring Asian longhorned beetles in Massachusetts
Melody Keena, Marie-José Côté, Phyllis S. Grinberg, William E. Wallner. 2011. Predicting the female flight capability of gypsy moths by using DNA markers
Jeremy R. deWaard, Andrew Mitchell, Melody Keena, David Gopurenko, Laura M. Boykin, Karen F. Armstrong, Michael G. Pogue, Joao Lima, Robin Floyd, Robert H. Hanner, Leland M. Humble. 2010. Towards a global barcode library for Lymantria (Lepidoptera: Lymantriinae) tussock moths of biosecurity concern
Melody Keena, Alice Vandel, Oldrich Pultar. 2010. Phenology of Lymantria monacha (Lepidoptera:Lymantriidae) laboratory reared on spruce foliage or a newly developed artificial diet
Melody Keena, Paul Moore. 2010. Effects of temperature on Anoplophora glabripennis (Coleoptera: Cerambycidae) larvae and pupae
M.E. Nehme, Melody Keena, A. Zhang, T.C. Baker, Z. Xu, K. Hoover. 2010. Evaluating the use of male-produced pheromone components and plant volatiles in two trap designs to monitor Anoplophora glabripennis
Maya Nehme, Melody Keena, Aijun Zhang, Kelli Hoover. 2009. Using Asian longhorned beetle male-produced pheromone and host volatiles for monitoring
Melody Keena, Juli Gould, Leah Bauer. 2009. Factors that influence emerald ash borer (Agrilus planipennis) adult longevity and oviposition under laboratory conditions
Nathan P. P. Havill, Michael Montgomery, Shigehiko Shiyake, Ashley Lamb, Melody Keena, Adalgisa Caccone. 2009. Hemlock woolly adelgid population genetics
Nathan P. P. Havill, Joanne Klein, Melody Keena, Adalgisa Caccone. 2009. Characterization of microsatellite loci for Laricobius nigrinus and L rubidus, predators of adelgids in North America
M.E. Nehme, Melody Keena, A. Zhang, T.C. Baker, K. Hoover. 2009. Attraction of Anoplophora glabripennis to male-produced phermonone and plant volatiles
Maya Nehme, Melody Keena, Aijun Zhang, Kelli Hoover. 2009. Field bioassays on the Asian longhorned beetle male-produced pheromone
Vicente Sanchez, Melody Keena, M.A. Keena. 2009. Allozyme Variation in Hemlock Woolly Adelgid (Hemiptera: Adelgidae) from the United States and China
Ryan P. Shanley, Melody Keena, Micheal M. Wheeler, Jarrod Leland, Ann E. Hajek. 2009. Evaluating the virulence and longevity of non-woven fiber bands impregnated with Metarhizium anisopliae against the Asian longhorned beetle, Anoplophora glabripennis (Coleoptera: Cerambycidae)
D.B. Roden, Robert A. A. Haack, Melody Keena, D.W. McKenney, F.D. Beall, P.M. Roden. 2009. Potential northern distribution of Asian longhorned beetle in North America
Melody Keena. 2009. Phenology of the Asian longhorned beetle under simulated annual environmental fluctuations
Brad Onken, Melody Keena. 2008. Hemlock woolly adelgid initiative: progress and future direction
Zhangjing Chen, Marshall S. White, Melody Keena, Therese M. M. Poland, Erin Clark. 2008. Evaluation of vacuum technology to kill larvae of the Asian longhorned beetle, Anoplophora glabripennis (Coleoptera: Cerambycidae), and the emerald ash borer, Agrilus planipennis (Coleoptera: Buprestidae), in wood
Maya Nehme, Kelli Hoover, Thomas Baker, Melody Keena, Aijun Zhang. 2008. Behavioral responses to Asian longhorned beetle pheromones
Melody Keena, Vicente Sanchez. 2008. How long does it take teneral adult Asian longhorned beetles to scleritize and then chew out of the wood?
Melody Keena, M.-J. Cote, Phyllis Grinberg, W.E. Wallner. 2008. World distribution of female flight and genetic variation in Lymantria dispar (Lepidoptera: Lymantriidae)
Melody Keena, Vicente Sanchez. 2007. Reproductive behaviors of Asian longhorned beetle
Melody Keena. 2007. Factors that influence Asian longhorned beetle pupation
Melody Keena, Vicente Sanchez. 2007. Reproductive behaviors of Anoplophora glabripennis (Coleoptera: Cerambycidae) in the laboratory
Melody Keena, Phyllis Grinberg, W.E. Wallner. 2007. Inheritance of female flight in Lymantria dispar (Lepidoptera: Lymantriidae)
Melody Keena. 2006. Effects of Temperature on Anoplophora glabripennis (Coleoptera: Cerambycidae) Adult Survival, Reproduction, and Egg Hatch
Melody Keena. 2005. Mating Parameters Associated with Fertility in Anoplophora Glabripennis (Coleoptera: Cerambycidae)
Alan N.D. Auclair, G. Fowler, A.T. M.K. Hogue, Melody Keena, D.R. Lance, R.M. McDowell, D.O. Oryang, A.J. Sawyer, A.J. Sawyer. 2005. Assessment of the Risk of Introduction of Anoplophora glabripennis in Municipal Solid Waste from the Quarantine Area of New York City to Landfills Outside of the Quarantine Area: a Pathway Analysis of the Risk of Spread and Establishment
Maureen E. Carter, E. Richard Hoebeke, Richard G. Harrison, Steven M. Bogdanowicz, Melody Keena, Alan Sawyer. 2005. Analysis of mtDNA Sequence Data for the Asian Longhorned Beetle (Anoplophora glabripennis): Evidence for Multiple Invasions in North America
David R. Gray, Melody Keena. 2005. A Comparison of Diapause in the Asian and North American Gypsy Moth? Implications for Population Establishment
Melody Keena. 2005. Effects of Timing of Larval Chill on Anoplophora glabripennis (Coleoptera: Cerambycidae) Survival and Pupation
Charlotte Nielsen, Melody Keena, Michael G. Milgroom, Ann E. Hajek. 2005. Asian Versus European Entomophaga maimaiga/Gypsy Moth Relations
Melody Keena, Melody Keena. 2005. Alternate Iron Sources for Use in Gypsy Moth Artificial Diet
Charlotte Nielsen, Melody Keena, Ann E. Hajek. 2005. Virulence and fitness of the fungal pathogen Entomophaga maimaiga in its host Lyrnantria dispar, for pathogen and host strains originating from Asia, Europe, and North America
Melody Keena. 2005. Pourable Artificial Diet for Rearing Anoplophora glabripennis (Coleoptera: Cerambycidae) and Methods to Optimize Larval Survival and Synchronize Development
Declan J. Fallon, Leellen F. Solter, Melody Keena, Michael McManus, James R. Cate, Lawrence M. Hanks. 2004. Susceptibility of Asian longhorned beetle, Anoplophora glabripennis (Motchulsky) (Coleoptera: Cerambycidae) to entomopathogenic nematodes
Melody Keena, Ann E. Hajek, Thomas L. M. Dubois, David R. Lance. 2003. Synthesis report on rearing Asian longhorned beetle
Melody Keena, Paul Moore, Steve M. Ulanecki. 2003. Effects of temperature on the life history parameters of Anoplophora Glabripennis (Coleoptera: Cerambycidae)
Michael T. Smith, Zhong-qi Yang, Franck Herard, Roger Fuester, Leah Bauer, Leellen Solter, Melody Keena, Vince D'Amico. 2003. Biological control of Anoplohora glabripennis Motsch.: a synthesis of current research programs
Melody Keena. 2003. Survival and Development of Lymantria monacha (Lepidoptera: Lymantriidae) on North American and Introduced Eurasian Tree Species
Melody Keena. 2002. Anoplophora glabripennis (Coleoptera: Cerambycidae) fecundity and longevity under laboratory conditions: comparison of populations from New York and Illinois on Acer saccharum
T.M. Withers, Melody Keena. 2001. Lymantria monacha (nun moth) and L. dispar (gypsy moth) survival and development on improved Pinus radiata
Melody Keena. 2001. Female Flight Propensity and Capability in Lymantria dispar (Lepidoptera: Lymantriidae) from Russia, North America, and Their Reciprocal F1 Hybrids
Melody Keena. 1996. Comparison of the Hatch of Lymantria dispar (Lepidoptera: Lymantriidae) Eggs from Russia and the United States After Exposure to Different Temperatures and Durations of Low Temperature
Melody Keena, Thomas M. ODell. 1994. Effects of laboratory testing on the gypsy moth (Lepidoptera:Lymantriidae)

University of California, Ph.D., Department of Entomology, 1988
University of California, M.S., Department of Entomology, 1985
University of California, B.S., Department of Entomology, 1983

Last updated March 10, 2026

Melody Keena, PhD

Treesearch Publications

Flight in Female Asian Spongy Moths

Ecology and Management of Invasive Species and Forest Ecosystems

Enhanced Mitigation and Rapid Response to Reduce Spread and Impact of Spotted Lanternfly in the United States

Biological Control of Hemlock Woolly Adelgid

Prescribed Fire as a Tool for Invasive Species Mitigation

Multimedia

Forestcast Season 1: Balance and Barrier - Episode 6: The Future of Forest Threats

Forestcast Season 1: Balance and Barrier - Episode 5: The Hornless Asian Longhorned Beetle