Patrick Tobin, PhD

Treesearch Publications

• Ashley N. Schulz, Nathan P. P. Havill, Travis D. Marsico, Matthew P. Ayres, Kamal J.K. Gandhi, Daniel A. Herms, Angela M. Hoover, Ruth A. Hufbauer, Andrew M. Liebhold, Kenneth F. Raffa, Kathryn A. Thomas, Patrick Tobin, Daniel R. Uden, Angela M. Mech. 2025. What is a specialist? Quantifying host breadth enables impact prediction for invasive herbivores
• Daniel R. Uden, Angela M. Mech, Nathan P. P. Havill, Ashley N. Schulz, Matthew P. Ayres, Daniel A. Herms, Angela M. Hoover, Kamal J. K. Gandhi, Ruth A. Hufbauer, Andrew M. Liebhold, Travis D. Marsico, Kenneth F. Raffa, Kathryn A. Thomas, Patrick Tobin, Craig R. Allen. 2022. Phylogenetic risk assessment is robust for forecasting the impact of European insects on North American conifers
• Rebecca Epanchin-Niell, Jieyi Lu, Alexandra Thompson, Patrick Tobin, David R. Gray, Andrew M. Liebhold. 2022. Socio-environmental drivers of establishment of Lymantria dispar, a nonnative forest pest, in the United States
• Ashley N. Schulz, Angela M. Mech, Matthew P. Ayres, Kamal J. K. Gandhi, Nathan P. P. Havill, Daniel A. Herms, Angela M. Hoover, Ruth A. Hufbauer, Andrew M. Liebhold, Travis D. Marsico, Kenneth F. Raffa, Patrick Tobin, Daniel R. Uden, Kathryn A. Thomas. 2021. Predicting non-native insect impact: focusing on the trees to see the forest
• Patrick Tobin, Brian Strom, Joseph A. Francese, Daniel A. Herms, Deborah G. McCullough, Therese M. M. Poland, Krista L. Ryall, Taylor Scarr, Peter J. Silk, Harold W. Thistle. 2021. Evaluation of Trapping Schemes to Detect Emerald Ash Borer (Coleoptera: Buprestidae)
• Jonathan A. Walter, Kristine L. Grayson, Laura Blackburn, Patrick Tobin, Derek M. Johnson. 2020. Spatiotemporal variability in Allee effects of invading gypsy moth populations
• Marissa A. Streifel, Patrick Tobin, Aubree M. Kees, Brian H. Aukema. 2019. Range expansion of Lymantria dispar dispar (L.) (Lepidoptera: Erebidae) along its north-western margin in North America despite low predicted climatic suitability
• Angela M. Mech, Kathryn A. Thomas, Travis D. Marsico, Daniel A. Herms, Craig R. Allen, Matthew P. Ayres, Kamal J. K. Gandhi, Jessica Gurevitch, Nathan P. Havill, Ruth A. Hufbauer, Andrew M. Liebhold, Kenneth F. Raffa, Ashley N. Schulz, Daniel R. Uden, Patrick Tobin. 2019. Evolutionary history predicts high‐impact invasions by herbivorous insects
• Stephanie Jagemann, Jennifer Juzwik, Patrick Tobin, Kenneth F. Raffa. 2018. Seasonal and Regional Distributions, Degree-Day Models, and Phoresy Rates of the Major Sap Beetle (Coleoptera: Nitidulidae) Vectors of the Oak Wilt Fungus, Bretziella fagacearum, in Wisconsin
• Patrick Tobin, Richard M. Turcotte, Laura Blackburn, John Juracko, Brian Simpson. 2017. The big chill: quantifying the effect of the 2014 North American cold wave on hemlock woolly adelgid populations in the central Appalachian Mountains
• Andrew M. Liebhold, Ludek Berec, Eckehard G. Brockerhoff, Rebecca S. Epanchin-Niell, Alan Hastings, Daniel A. Herms, John M. Kean, Deborah G. McCullough, David M. Suckling, Patrick Tobin, Takehiko Yamanaka. 2016. Eradication of Invading Insect Populations: From Concepts to Applications
• Ann E. Hajek, Patrick Tobin, Kyle J. Haynes. 2015. Replacement of a dominant viral pathogen by a fungal pathogen does not alter the collapse of a regional forest insect outbreak
• Samita Limbu, Katie Cassidy, Melody Keena, Patrick Tobin, Kelli Hoover. 2015. Host range specificity of Scymnus camptodromus (Coleoptera: Coccinellidae), a predator of hemlock woolly adelgid (Hemiptera: Adelgidae)
• Patrick Tobin, Andrew M. Liebhold, E. Anderson Roberts, Laura Blackburn. 2015. Estimating spread rates of non-native species: the gypsy moth as a case study
• Kevin M. Bigsby, Mark J. Ambrose, Patrick Tobin, Erin O. Sills. 2014. The cost of gypsy moth sex in the city
• Patrick Tobin, Joshua L. Bolyard, Ksenia S. Onufrieva, Andrea D. Hickman. 2014. The Effect of Male and Female Age on Lymantria dispar (Lepidoptera: Lymantriidae) Fecundity
• Patrick Tobin, Laura Blackburn, Rebecca H. Gray, Christopher T. Lettau, Andrew M. Liebhold, Kenneth F. Raffa. 2013. Using delimiting surveys to characterize the spatiotemporal dynamics facilitates the management of an invasive non-native insect
• Christopher M. Ranger, Patrick Tobin, Michael E. Reding, Alicia M. Bray, Jason B. Oliver, Peter B. Schultz, Steven D. Frank, Anand B. Persad. 2013. Interruption of the Semiochemical-Based Attraction of Ambrosia Beetles to Ethanol-Baited Traps and Ethanol-Injected Trap Trees by Verbenone
• Patrick Tobin, Richard M. Turcotte, Daniel A. Snider. 2013. When one is not necessarily a lonely number: initial colonization dynamics of Adelges tsugae on eastern hemlock, Tsuga canadensis
• K.L. Frank, Patrick Tobin, Jr. Thistle, Laurence S. Kalkstein. 2013. Interpretation of gypsy moth frontal advance using meteorology in a conditional algorithm
• Patrick Tobin, Ksenia S. Onufrieva, Kevin W. Thorpe. 2012. The relationship between male moth density and female mating success in invading populations of Lymantria dispar
• Richard M. Turcotte, Thomas R. Elliott, Mary Ann Fajvan, Yong-Lak Park, Daniel A. Snider, Patrick Tobin. 2012. Effects of ice storm damage on hardwood survival and growth in Ohio
• Patrick Tobin, Ann E. Hajek. 2012. Release, establishment, and initial spread of the fungal pathogen Entomophaga maimaiga in island populations of Lymantria dispar
• Patrick Tobin, Barry B. Bai, Donald A. Eggen, Donna Leonard. 2012. The ecology, geopolitics, and economics of managing Lymantria dispar in the United States
• David Maxwell Suckling, Patrick Tobin, Deborah G. McCullough, Daniel A. Herms. 2012. Combining tactics to exploit allee effects for eradication of alien insect populations
• Laura Blackburn, Donna S. Leonard, Patrick Tobin. 2011. The use of Bacillus thuringiensis kurstaki for managing gypsy moth populations under the Slow the Spread Program, 1996-2010, relative to the distributional range of threatened and endangered species
• Ann E. Hajek, Patrick Tobin. 2011. Introduced pathogens follow the invasion front of a spreading alien host
• Patrick Tobin, Aijun Zhang, Ksenia Onufrieva, Donna Leonard. 2011. Field evaluation of effect of temperature on release of Disparlure from a pheromone-baited trapping system used to monitor gypsy moth (Lepidoptera: Lymantriidae)
• Kevin M. Bigsby, Patrick Tobin, Erin O. Sills. 2011. Anthropogenic drivers of gypsy moth spread
• Shi Chen, Shelby J. Fleischer, Patrick Tobin, Michael C. Saunders. 2011. Projecting insect voltinism under high and low greenhouse gas emission conditions
• Patrick Tobin, Ludek Berec, Andrew M. Liebhold. 2011. Exploiting Allee effects for managing biological invasions
• Patrick Tobin, Julie Van Stappen, Laura Blackburn. 2011. Visitation rates to the Apostle Islands National Lakeshore and the introduction of the non-native species Lymantria dispar (L.)
• Patrick Tobin, Andrea Diss-Torrance, Laura Blackburn, Brian D. Brown. 2011. What does "local" firewood buy you? Managing the risk of invasive species introduction
• E. Vercken, A.M. Kramer, Patrick Tobin, J.M. Drake. 2011. Critical patch size generated by Allee effect in gypsy moth, Lymantria dispar (L.)
• Patrick Tobin, Andrea Diss-Torrance, Laura Blackburn, Brian D. Brown. 2010. What does "local" firewood buy you? Managing the risk of invasive species introduction
• Jody Timer, Patrick Tobin, Michael C. Saunders. 2010. Geographic variation in diapause induction: The grape berry moth (Lepidoptera: Tortricidae)
• Ann E. Hajek, Patrick Tobin. 2010. Micro-managing arthropod invasions: eradication and control of invasive arthropods with microbes
• Patrick Tobin, Julie Van Stappen, Laura Blackburn. 2010. Human visitation rates to the Apostle Islands National Lakeshore and the introduction of the non-native species Lymantria dispar (L.)
• Andrew M. Liebhold, Patrick Tobin. 2010. Exploiting the Achilles heels of pest invasions: allee effects, stratified dispersal and management of forest insect establishment and spread
• Patrick Tobin, Kenneth T. Klein, Donna S. Leonard. 2009. Gypsy moth (Lepidoptera: Lymantriidae) flight behavior and phenology based on field-deployed automated pheromone-baited traps
• Ann E. Hajek, Patrick Tobin. 2009. A space-time odyssey: movement of gypsy moth and its pathogens
• Patrick Tobin, James L. Frazier. 2009. A slide down a slippery slope: ethical guidelines in the dissemination of computer-based presentations
• Kenneth F. Raffa, Brian Aukema, Barbara J. Bentz, Allan Carroll, Nadir Erbilgin, Daniel A. Herms, Jeffrey A. Hicke, Richard W. Hofstetter, Steven Katovich, B. Staffan Lindgren, Jesse Logan, William Mattson, A. Steven Munson, Daniel J. Robison, Diana L. Six, Patrick Tobin, Philip A. Townsend, Kimberly F. Wallin. 2009. A literal use of "forest health" safeguards against misuse and misapplication
• Patrick Tobin, Christelle Robinet, Derek M. Johnson, Stefanie L. Whitmire, Ottar N. Bjornstad, Andrew M. Liebhold. 2009. The role of Allee effects in gypsy moth, Lymantria dispar (L.), invasions
• Patrick Tobin, Ann E. Hajek, Laura Blackburn, Joshua J. Hannam, Andrea Diss-Torrance, Kenneth F. Raffa, Charlotte Nielson. 2009. Space-time interactions between gypsy moth and associated entomopathogens in newly established populations
• Ann E. Hajek, Patrick Tobin, Laura Blackburn, Joshua J. Hannam, Andrea Diss-Torrance, Kenneth Raffa, Charlotte Nielsen. 2009. Dispersal of gypsy moth pathogens to newly established host populations
• Mark Ambrose, Simone Bauch, Kevin Bigsby, Yun Wu, Patrick Tobin, Erin Sills. 2009. Gypsy moth sex in the city: estimating costs and losses
• Rebecca Hoffman Gray, Craig G. Lorimer, Patrick Tobin, Kenneth F. Raffa. 2008. Preoutbreak dynamics of a recently established invasive herbivore: roles of natural enemies and habitat structure in stage-specific performance of gypsy moth (Lepidoptera: Lymantriidae) populations in northeastern Wisconsin
• Patrick Tobin, Stephanie L. Whitmire, Derek M. Johnson, Ottar N. Bjornstad, Andrew M. Liebhold. 2008. Allee effects and gypsy moth invasions
• Richard M. Turcotte, Patrick Tobin. 2008. Relationship between the fungal pathogen Entomophaga maimaiga and the gypsy moth on the Monongahela National Forest
• Michael T. Smith, Patrick Tobin, Jinquan Wu, Weizhi He, Xuenong Xu, Gerhard Gries, Regine Gries, John H. Borden, Jean J. Turgeon, Peter de Groot. 2008. Behavioral ecology of host selection in the Asian longhorned beetle: implications for surveying, detecting, and monitoring adult beetles
• Ann E. Hajek, Joshua Hannam, Charlotte Nielsen, Andrea Diss-Torrance, Ken Raffa, Patrick Tobin. 2008. Dispersal of gypsy moth pathogens into areas newly colonized by gypsy moth
• Patrick Tobin. 2008. Cost analysis and biological ramifications for implementing the gypsy moth Slow the Spread Program
• Andrew M. Liebhold, Patrick Tobin. 2008. Population ecology of insect invasions and their management
• Patrick Tobin, Laura Blackburn. 2008. Long-distance dispersal of the gypsy moth (Lepidoptera: Lymantriidae) facilitated its initial invasion of Wisconsin
• Patrick Tobin, Sudha Nagarkatti, Greg Loeb, Michael C. Saunders. 2008. Historical and projected interactions between climate change and insect voltinism in a multivoltine species
• Kevin W. Thorpe, Ksenia S. Tcheslavskaia, Patrick Tobin, Laura Blackburn, Donna S. Leonard, E. Anderson Roberts. 2007. Persistent effects of aerial applications of disparlure on gypsy moth: trap catch and mating success
• Randall Morin, Andrew M. Liebhold, Patrick Tobin, Kurt W. Gottschalk, Eugene Luzader. 2007. Spread of beech bark disease in the eastern United States and its relationship to regional forest composition
• Patrick Tobin, Laura Blackburn. 2007. Slow the Spread: a national program to manage the gypsy moth
• Patrick Tobin, Kevin W. Thorpe, Laura Blackburn. 2007. Multi-year evaluation of mating disruption treatments against gypsy moth
• Patrick Tobin. 2007. Space--time patterns during the establishmentof a nonindigenous species
• Patrick Tobin, Andrew M. Liebhold, E. Anderson Roberts. 2007. Comparison of methods for estimating the spread of a non-indigenous species
• Patrick Tobin, Stephanie L. Whitmire, Derek M. Johnson, Ottar N. Bjornstad, Andrew M. Liebhold. 2007. Invasion speed is affected by geographical variation in the strength of Allee effects
• Derk M. Johnson, Andrew M. Liebhold, Patrick Tobin, Ottar N. Bjornstad. 2006. Allee effects and pulsed invasion by the gypsy moth
• Stefanie L. Whitmire, Patrick Tobin. 2006. Persistence of invading gypsy moth populations in the United States
• Andrew M. Liebhold, Patrick Tobin. 2006. Growth of newly established alien populations: comparison of North American gypsy moth colonies with invasion theory
• Cynthia Huebner, Patrick Tobin. 2006. Invasibility of mature and 15-year-old deciduous forests by exotic plants
• Patrick Tobin. 2005. The Spread of Gypsy Moth in Wisconsin: a New Paradigm for the Midwest Invasion?
• Patrick Tobin, Ottar N. Bjornstad. 2005. Roles of dispersal, stochasticity, and nonlinear dynamics in the spatial structuring of seasonal natural enemy-victim populations
• Patrick Tobin, Stefanie L. Whitmire. 2005. Spread of Gypsy Moth (Lepidoptera: Lymantriidae) and Its Relationship to Defoliation
• Patrick Tobin, Stefanie L. Whitmire. 2005. Persistence of Invading Gypsy Moth Populations [abstract]
• Patrick Tobin, Michael T. Smith, Guohong Li, Ruitong Gao. 2005. Dispersal and Spatiotemporal Dynamics of Asian Longhorned Beetle in China
• Patrick Tobin. 2004. Estimation of the spatial autocorrelation function: consequences of sampling dynamic populations in space and time
• Patrick Tobin, Alexei A. Sharov, Andrew M. Liebhold, Donna S. Leonard, E. Anderson Roberts, Matthew R. Learn. 2004. Management of the gypsy moth through a decision algorithm under the STS project

• Penn State University, University Park, PA, Ph.D., Entomology with minors in Statistics and Operations Research, 2002
• Penn State University, University Park, PA., M.S., Entomology, 1997
• University of Georgia, Athens, GA., B.S., Environmental Health Sciences, 1991