Samuel A. Cushman, PhD

Treesearch Publications

• Ho Yi Wan, Michael A. Lommler, Samuel A. Cushman, Jamie S. Sanderlin, Joseph L. Ganey, Andrew J. Sanchez Meador, Paul Beier. 2025. A multi-level, multi-scale comparison of LiDAR- and LANDSAT-based habitat selection models of Mexican spotted owls in a post-fire landscape
• Laura Bosco, Valentin Moser, Mirkka M. Jones, Oystein Opedal, Otso Ovaskainen, Gerber Sonja, Roel Van Klink, Samuel A. Cushman, Raphael Arlettaz, Alain Jacot. 2023. Habitat area and local habitat conditions outweigh fragmentation effects on insect communities in vineyards
• Kelsey E. Paolini, Michael K. Schwartz, Megan M. Friggens, Samuel A. Cushman, Julie Weckworth, Joseph D. Holbrook. 2023. A review of population and landscape level dynamics associated with pneumonia outbreaks in bighorn sheep with implications for land management
• Thomas E. Dilts, Katherine Zeller, Samuel A. Cushman, Eveline S. Larrucea, Miranda M. Crowell, Nathan W. Byer, Kevin T. Shoemaker, Marjorie D. Matocq. 2023. Pygmy rabbit habitat network reveals threats and opportunities for management and conservation
• Siddharth Unnithan Kumar, Samuel A. Cushman. 2022. Connectivity modelling in conservation science: A comparative evaluation
• Logan Hysen, Danial Nayeri, Samuel A. Cushman, Ho Yi. Wan. 2022. Background sampling for multi-scale ensemble habitat selection modeling: Does the number of points matter?
• Gavin M. Jones, Andrew J. Shirk, Zhiqiang Yang, Raymond J. Davis, Joseph L. Ganey, R. J. Gutierrez, Sean P. Healey, Shaula J. Hedwall, Serra J. Hoagland, Ron Maes, Karl Malcolm, Kevin S. McKelvey, Jamie S. Sanderlin, Michael K. Schwartz, Mark E. Seamans, Ho Yi Wan, Samuel A. Cushman. 2022. Spatial and temporal dynamics of Mexican spotted owl habitat in the southwestern US
• Andrew J. Shirk, Gavin M. Jones, Zhiqiang Yang, Raymond J. Davis, Joseph L. Ganey, R. J. Gutierrez, Sean P. Healey, Shaula J. Hedwall, Serra J. Hoagland, Ron Maes, Karl Malcolm, Kevin S. McKelvey, Carly Vynne, Jamie S. Sanderlin, Michael K. Schwartz, Mark E. Seamans, Ho Yi Wan, Samuel A. Cushman. 2022. Automated habitat monitoring systems linked to adaptive management: A new paradigm for species conservation in an era of rapid environmental change
• Eric Ash, Samuel A. Cushman, David W. Macdonald, Tim Redford, Zaneta Kaszta. 2022. How important are resistance, dispersal ability, population density and mortality in temporally dynamic simulations of population connectivity? A case study of tigers in southeast Asia
• Jay M. Schoen, Amrita Neelakantan, Samuel A. Cushman, Trishna Dutta, Bilal Habib, Yadvendradev V Jhala, Indranil Mondal, Uma Ramakrishnan, P. Anuradha Reddy, Swati Saini, Sandeep Sharma, Prachi Thatte, Bibek Yumnam, Ruth DeFries. 2022. Synthesizing habitat connectivity analyses of a globally important human-dominated tiger-conservation landscape
• Siddharth Unnithan Kumar, Jonathon Turnbull, Oscar Hartman Davies, Timothy Hodgetts, Samuel A. Cushman. 2022. Moving beyond landscape resistance: Considerations for the future of connectivity modelling and conservation science
• Changxiu Cheng, Samuel A. Cushman, Hung-Chak Ho, Peichao Gao. 2022. Geographic complexity: Concepts, theories, and practices
• L. Chiaverini, D. W. Macdonald, H. M. Bothwell, A. J. Hearn, S. M. Cheyne, I. Haidir, L. T. B. Hunter, Z. Kaszta, E. A. Macdonald, J. Ross, Samuel A. Cushman. 2022. Multi-scale, multivariate community models improve designation of biodiversity hotspots in the Sunda Islands
• S. A. Dar, S. K. Singh, H. Y. Wan, Samuel A. Cushman, T. Bashir, S. Sathyakumar. 2022. Future land use and climate change escalate connectivity loss for Himalayan brown bears
• Mihai-Sorin Stupariu, Samuel A. Cushman, Alin-Ionut Plesoianu, Ileana Patru-Stupariu, Christine Furst. 2022. Machine learning in landscape ecological analysis: A review of recent approaches
• Gavin M. Jones, J. Ayars, Sean A. Parks, Helen E. Chmura, Samuel A. Cushman, Jamie S. Sanderlin. 2022. Pyrodiversity in a warming world: Research challenges and opportunities
• Ugyen Penjor, Samuel A. Cushman, Zaneta M. Kaszta, Sherub Sherub, David W. Macdonald. 2022. Effects of land use and climate change on functional and phylogenetic diversity of terrestrial vertebrates in a Himalayan biodiversity hotspot
• Ugyen Penjor, Christos Astaras, Samuel A. Cushman, Zaneta Kaszta, David W. Macdonald. 2022. Contrasting effects of human settlement on the interaction among sympatric apex carnivores
• Davis E. Blasini, Dan F. Koepke, Kevin C. Grady, Gerard J. Allan, Catherine A. Gehring, Thomas G. Whitham, Samuel A. Cushman, Kevin R. Hultine. 2021. Adaptive trait syndromes along multiple economic spectra define cold and warm adapted ecotypes in a widely distributed foundation tree species
• Pyae Phyoe Kyaw, David W. Macdonald, Ugyen Penjor, Saw Htun, Hla Naing, Dawn Burnham, Zaneta Kaszta, Samuel A. Cushman. 2021. Investigating carnivore guild structure: Spatial and temporal relationships amongst threatened felids in Myanmar
• Luciano Atzeni, Samuel A. Cushman, Jun Wang, Philip Riordan, Kun Shi, David Bauman. 2021. Evidence of spatial genetic structure in a snow leopard population from Gansu, China
• Samuel A. Cushman. 2021. Entropy in landscape ecology: A quantitative textual multivariate review
• Samuel A. Cushman. 2021. Thermodynamic consistency of the Cushman method of computing the configurational entropy of a landscape lattice
• Samuel A. Cushman. 2021. Generalizing Boltzmann configurational entropy to surfaces, point patterns and landscape mosaics
• Andrew J. Shirk, Erin L. Landguth, Samuel A. Cushman. 2021. The effect of gene flow from unsampled demes in landscape genetic analysis
• Eric Ash, David W. Macdonald, Samuel A. Cushman, Adisorn Noochdumrong, Tim Redford, Żaneta Kaszta. 2021. Optimization of spatial scale, but not functional shape, affects the performance of habitat suitability models: a case study of tigers (Panthera tigris) in Thailand
• Luca Chiaverini, Ho Yi Wan, Beth Hahn, Amy Cilimburg, Tzeidle N. Wasserman, Samuel A. Cushman. 2021. Effects of non-representative sampling design on multi-scale habitat models: Flammulated owls in the Rocky Mountains.
• S. A. Dar, S. K. Singh, H. Y. Wan, V. Kumar, Samuel A. Cushman, S. Sathyakumar. 2021. Projected climate change threatens Himalayan brown bear habitat more than human land use
• Violeta Lopez‐Marquez, Samuel A. Cushman, Jose Templado, Ho Yi Wan, Helen M. Bothwell, Annie Machordom. 2021. Genetic connectivity of two marine gastropods in the Mediterranean Sea: Seascape genetics reveals species‐specific oceanographic drivers of gene flow
• Michael K. Lucid, Ho Yi Wan, Shannon Ehlers, Lacy Robinson, Leona K. Svancara, Andrew Shirk, Samuel A. Cushman. 2021. Land snail microclimate niches identify suitable areas for climate refugia management on a montane landscape
• Katherine Zeller, Cody A. Schroeder, Ho Yi Wan, Gail Collins, Kristin Denryter, Andrew F. Jakes, Samuel A. Cushman. 2021. Forecasting habitat and connectivity for pronghorn across the Great Basin ecoregion
• Ugyen Penjor, Zaneta Kaszta, David W. Macdonald, Samuel A. Cushman. 2021. Prioritizing areas for conservation outside the existing protected area network in Bhutan: the use of multi-species, multi-scale habitat suitability models
• A. Mohammadi, K. Almasieh, D. Nayeri, F. Ataei, A. Khani, J. V López-Bao, V. Penteriani, Samuel A. Cushman. 2021. Identifying priority core habitats and corridors for effective conservation of brown bears in Iran
• Alireza Mohammadi, Kamran Almasieh, Ho Yi Wan, Danial Nayeri, Amir Alambeigi, Jason I Ransom, Samuel A. Cushman. 2021. Integrating spatial analysis and questionnaire survey to better understand human-onager conflict in Southern Iran
• Mitra Menon, Justin C. Bagley, Gerald F. M. Page, Amy V Whipple, Anna W. Schoettle, Christopher J. Still, Christian Wehenkel, Kristen M. Waring, Lluvia Flores-Renteria, Samuel A. Cushman, Andrew J. Eckert. 2021. Adaptive evolution in a conifer hybrid zone is driven by a mosaic of recently introgressed and background genetic variants
• Katherine Zeller, Samuel A. Cushman, Nicholas J. Van Lanen, John D. Boone, Elisabeth Ammon. 2021. Targeting conifer removal to create an even playing field for birds in the Great Basin
• Caroline Charao Sartor, Samuel A. Cushman, Ho Yi Wan, Rafael Kretschmer, Javier A. Pereira, Nadia Bou, Mariana Cosse, Susana Gonzalez, Eduardo Eizirik, Thales Renato O. de Freitas, Tatiane Campos. Trigo. 2021. The role of the environment in the spatial dynamics of an extensive hybrid zone between two neotropical cats
• Guilherme Costa Alvarenga, Luca Chiaverini, Samuel A. Cushman, Egil Droge, David W. Macdonald, Daniel Luis Zanella Kantek, Ronaldo Goncalves Morato, Jeffrey J. Thompson, Ricardo B. L. M. Oscar, Leandro Abade, Fernando Cesar Cascelli de Azevedo, Emiliano E. Ramalho, Zaneta Kaszta. 2021. Multi-scale path-level analysis of jaguar habitat use in the Pantanal ecosystem
• Deborah M. Finch, Jack L. Butler, Justin B. Runyon, Christopher J. Fettig, Francis F. Kilkenny, Shibu Jose, Susan J. Frankel, Samuel A. Cushman, Richard C. Cobb, Jeffrey S. Dukes, Jeffrey A. Hicke, Sybill K. Amelon. 2021. Effects of Climate Change on Invasive Species
• Samuel A. Cushman. 2020. Location, location, location - scale, scale, scale: Mexican spotted owl habitat
• Sylvia Kantor, Samuel A. Cushman, Joe Ganey, Gavin M. Jones, Ho Yi Wan, Karl Malcolm, Shaula Hedwall. 2020. Through the smoke: spotted owls, wildfire, and forest restoration
• Z. Kaszta, Samuel A. Cushman, D. W. Macdonald. 2020. Prioritizing habitat core areas and corridors for a large carnivore across its range
• Zaneta Kaszta, Samuel A. Cushman, Saw Htun, Hla Naing, Dawn Burnham, David W. Macdonald. 2020. Simulating the impact of Belt and Road initiative and other major developments in Myanmar on an ambassador felid, the clouded leopard, Neofelis nebulosa
• Lin-Feng Li, Samuel A. Cushman, Yan-Xia He, Yong Li. 2020. Genome sequencing and population genomics modeling provide insights into the local adaptation of weeping forsythia
• Mohsen Ahmadi, Mohammad S. Farhadinia, Samuel A. Cushman, Mahmoud-Reza Hemami, Bagher Nezami Balouchi, Houman Jowkar, David W. Macdonald. 2020. Species and space: A combined gap analysis to guide management planning of conservation areas
• Mohammad Reza Ashrafzadeh, Rasoul Khosravi, Mohammad Ali Adibi, Atie Taktehrani, Ho Yi Wan, Samuel A. Cushman. 2020. A multi-scale, multi-species approach for assessing effectiveness of habitat and connectivity conservation for endangered felids
• Luciano Atzeni, Samuel A. Cushman, Defeng Bai, Jun Wang, Pengju Chen, Kun Shi, Philip Riordan. 2020. Meta-replication, sampling bias, and multi-scale model selection: A case study on snow leopard (Panthera uncia) in western China
• Zuzana Burivalova, Edward T. Game, Bambang Wahyudi, Ruslandi, Mohamad Rifqi, Ewan MacDonald, Samuel A. Cushman, Maria Voigt, Serge Wich, David S. Wilcove. 2020. Does biodiversity benefit when the logging stops? An analysis of conservation risks and opportunities in active versus inactive logging concessions in Borneo
• Sarah K. Carter, David S. Pilliod, Travis Haby, Karen L. Prentice, Cameron L. Aldridge, Patrick J. Anderson, Zachary H. Bowen, John B. Bradford, Samuel A. Cushman, Joseph C. DeVivo, Michael C. Duniway, Ryan S. Hathaway, Lisa Nelson, Courtney A. Schultz, Rudy M. Schuster, E. Jamie Trammell, Jake F. Weltzin. 2020. Bridging the research-management gap: Landscape science in practice on public lands in the western United States
• Milena F. Diniz, Samuel A. Cushman, Ricardo B. Machado, Paulo De Marco Junior. 2020. Landscape connectivity modeling from the perspective of animal dispersal
• Peichao Gao, Haoyu Wang, Samuel A. Cushman, Changxiu Cheng, Changqing Song, Sijing Ye. 2020. Sustainable land-use optimization using NSGA-II: Theoretical and experimental comparisons of improved algorithms
• Kristen Waring, Samuel A. Cushman, Andrew Eckert, Lluvia Flores-Renteria, Richard Sniezko, Christopher Still, Christian Wehenkel, Amy Whipple, Michael Wing, Justin Bagley, Ethan Bucholz, Socorro Gonzalez-Elizondo, Marja Haagsma, Jessica Hartsell, Cory Garms, Evan Heck, Jeremy Johnson, Erin Landguth, Alejandro Leal Saenz, Mitra Menon, Ehren Moler, Gerald Page, John D. Shaw, Andrew Shirk, Jared Swenson. 2020. Ecology and evolution of a tree species challenged by dual threats
• Kristen Waring, Samuel A. Cushman, Andrew Eckert, Lluvia Flores, Richard Sniezko, Christopher Still, Christian Wehenkel, Amy Whipple, Michael Wing, Justin Bagley, Emily Boes, Ethan Bucholz, Jonathan Burnett, Antonio Castilla, Jessica Dabell, Chris Friedline, Cory Garms, Evan Heck, Jeremy Johnson, Erin Landguth, Alejandro Leal Sáenz, Briana Mcteague, Mitra Menon, Ehren Moler, Gerald Page, Andrew Shirk. 2020. Collaborative Research for Sustainable Management of Southwestern White Pine
• Gavin M. Jones, R. J. Gutierrez, William M. Block, Peter C. Carlson, Emily J. Comfort, Samuel A. Cushman, Raymond J. Davis, Stephanie A. Eyes, Alan B. Franklin, Joseph L. Ganey, Shaula Hedwall, John J. Keane, Rodd Kelsey, Damon B. Lesmeister, Malcolm P. North, Susan L. Roberts, Jeremy T. Rockweit, Jamie S. Sanderlin, Sarah C. Sawyer, Ben Solvesky, Douglas J. Tempel, Ho Yiwan, A. Leroy Westerling, Gary C. White, M. Zachariah Peery. 2020. Spotted owls and forest fire: Comment
• Glenn T. Howe, Keith Jayawickrama, Scott E. Kolpak, Jennifer Kling, Matt Trappe, Valerie Hipkins, Terrance Ye, Stephanie Guida, Richard Cronn, Samuel A. Cushman, Susan McEvoy. 2020. An Axiom SNP genotyping array for Douglas-fir
• Jeanne C. Chambers, Craig R. Allen, Samuel A. Cushman. 2020. Editorial: Operationalizing the concepts of resilience and resistance for managing ecosystems and species at risk
• Jeanne C. Chambers, Craig R. Allen, Samuel A. Cushman. 2020. Operationalizing the concepts of resilience and resistance for managing ecosystems and species at risk
• Corina Maurer, Laura Bosco, Elisabeth Klaus, Samuel A. Cushman, Raphael Arlettaz, Alain Jacot. 2020. Habitat amount mediates the effect of fragmentation on a pollinator's reproductive performance, but not on its foraging behaviour
• Ho Yi Wan, Samuel A. Cushman, Joseph L. Ganey. 2020. The effect of scale in quantifying fire impacts on species habitats
• Roya Adavodi, Rasoul Khosravi, Samuel A. Cushman, Mohammad Kaboli. 2019. Topographical features and forest cover influence landscape connectivity and gene flow of the Caucasian pit viper, Gloydius caucasicus (Nikolsky, 1916), in Iran
• Samuel A. Cushman, Kevin McGarigal. 2019. Metrics and models for quantifying ecological resilience at landscape scales
• P. A. Reddy, J.-P Puyravaud, Samuel A. Cushman, H. Segu. 2019. Spatial variation in the response of tiger gene flow to landscape features and limiting factors
• Michael K. Lucid, Andrew Rankin, Jack Sullivan, Lacy Robinson, Shannon Ehlers, Samuel A. Cushman. 2019. A carnivores’ oasis? An isolated fisher (Pekania pennanti) population provides insight on persistence of a metapopulation
• Laura Bosco, Ho Yi Wan, Samuel A. Cushman, Raphael Arlettaz, Alain Jacot. 2019. Separating the effects of habitat amount and fragmentation on invertebrate abundance using a multi-scale framework
• Andrew J. Hearn, Samuel A. Cushman, Benoit Goossens, Joanna Ross, Ewan A. Macdonald, Luke T. B. Hunter, David W. Macdonald. 2019. Predicting connectivity, population size and genetic diversity of Sunda clouded leopards across Sabah, Borneo
• David W. Macdonald, Helen M. Bothwell, Zaneta Kaszta, Eric Ash, Gilmoore Bolongon, Dawn Burnham, Ozgun Emre Can, Ahimsa Campos‐Arceiz, Phan Channa, Gopalasamy Reuben Clements, Andrew J. Hearn, Laurie Hedges, Saw Htun, Jan F. Kamler, Kae Kawanishi, Ewan A. Macdonald, Shariff Wan Mohamad, Jonathan Moore, Hla Naing, Manabu Onuma, Ugyen Penjor, Akchousanh Rasphone, Darmaraj Mark Rayan, Joanna Ross, Priya Singh, Cedric Kai Wei Tan, Jamie Wadey, Bhupendra P. Yadav, Samuel A. Cushman. 2019. Multi‐scale habitat modelling identifies spatial conservation priorities for mainland clouded leopards (Neofelis nebulosa)
• Rasoul Khosravi, Mahmoud-Reza Hemami, Samuel A. Cushman. 2019. Multi-scale niche modeling of three sympatric felids of conservation importance in central Iran
• Mitra Menon, Erin Landguth, Alejandro Leal‐Saenz, Justin C. Bagley, Anna W. Schoettle, Christian Wehenkel, Lluvia Flores‐Renteria, Samuel A. Cushman, Kristen M. Waring, Andrew J. Eckert. 2019. Tracing the footprints of a moving hybrid zone under a demographic history of speciation with gene flow
• Jeanne C. Chambers, Craig R. Allen, Samuel A. Cushman. 2019. Operationalizing ecological resilience concepts for managing species and ecosystems at risk
• Gilda Shahnaseri, Mahmoud-Reza Hemami, Rasoul Khosravi, Shima Malakoutikhah, Maryam Omidi, Samuel A. Cushman. 2019. Contrasting use of habitat, landscape elements, and corridors by grey wolf and golden jackal in central Iran
• Tania Barros, Joao Carvalho, Carlos Fonseca, Samuel A. Cushman. 2019. Assessing the complex relationship between landscape, gene flow, and range expansion of a Mediterranean carnivore
• Zaneta Kaszta, Samuel A. Cushman, Andrew J. Hearn, Dawn Burnham, Ewan A. Macdonald, Benoit Goossens, Senthilvel K. S. S. Nathan, David W. Macdonald. 2019. Integrating Sunda clouded leopard (Neofelis diardi) conservation into development and restoration planning in Sabah (Borneo)
• Ho Yi Wan, Samuel A. Cushman, Joseph L. Ganey. 2019. Recent and projected future wildfire trends across the ranges of three spotted owl subspecies under climate change
• Ho Yi Wan, Samuel A. Cushman, Joseph L. Ganey. 2019. Improving habitat and connectivity model predictions with multi-scale resource selection functions from two geographic areas
• Katherine Zeller, Megan K. Jennings, T. Winston Vickers, Holly B. Ernest, Samuel A. Cushman, Walter M. Boyce. 2018. Are all data types and connectivity models created equal? Validating common connectivity approaches with dispersal data
• Jeremy S. Littell, Donald McKenzie, Ho Yi Wan, Samuel A. Cushman. 2018. Climate change and future wildfire in the Western United States: an ecological approach to nonstationarity
• Kevin McGarigal, Maritza Mallek, Becky Estes, Marilyn Tierney, Terri Walsh, Travis Thane, Hugh Safford, Samuel A. Cushman. 2018. Modeling historical range of variability and alternative management scenarios in the upper Yuba River watershed, Tahoe National Forest, California
• Michael K. Lucid, Shannon Ehlers, Lacy Robinson, Samuel A. Cushman. 2018. Beer, brains, and brawn as tools to describe terrestrial gastropod species richness on a montane landscape
• Samuel A. Cushman, Tzeidle N. Wasserman. 2018. Landscape applications of machine learning: Comparing random forests and logistic regression in multi-scale optimized predictive modeling of American marten occurrence in northern Idaho, USA [Chapter 9]
• Andrew J. Shirk, Samuel A. Cushman, Kristen M. Waring, Christian A. Wehenkel, Alejandro Leal-Saenz, Chris Toney, Carlos A. Lopez-Sanchez. 2018. Southwestern white pine (Pinus strobiformis) species distribution models project a large range shift and contraction due to regional climatic changes
• Robert E. Keane, Samuel A. Cushman. 2018. Best friends forever: The whitebark pine and Clark's nutcracker
• Ho Yi Wan, Samuel A. Cushman, Joseph L. Ganey. 2018. Habitat fragmentation reduces genetic diversity and connectivity of the Mexican spotted owl: A simulation study using empirical resistance models
• Ewan A. Macdonald, Samuel A. Cushman, Erin L. Landguth, Andrew J. Hearn, Yadvinder Malhi, David W. Macdonald. 2018. Simulating impacts of rapid forest loss on population size, connectivity and genetic diversity of Sunda clouded leopards (Neofelis diardi) in Borneo
• Ho Yi Wan, Joseph L. Ganey, Christina D. Vojta, Samuel A. Cushman. 2018. Managing emerging threats to spotted owls
• Megan M. Friggens, Mary I. Williams, Karen Bagne, Tosha T. Wixom, Samuel A. Cushman. 2018. Effects of climate change on terrestrial animals [Chapter 9]
• Zaneta Kaszta, Samuel A. Cushman, Claudio Sillero-Zubiri, Eleonore Wolff, Jorgelina Marino. 2018. Where buffalo and cattle meet: Modelling interspecific contact risk using cumulative resistant kernels
• Mitra Menon, Justin C. Bagley, Christopher J. Friedline, Amy V. Whipple, Anna W. Schoettle, Alejandro Leal-Saenz, Christian Wehenkel, Francisco Molina-Freaner, Lluvia Flores-Renteria, M. Socorro Gonzalez-Elizondo, Richard A. Sniezko, Samuel A. Cushman, Kristen M. Waring, Andrew J. Eckert. 2017. The role of hybridization during ecological divergence of southwestern white pine (Pinus strobiformis) and limber pine (P. flexilis)
• Joseph L. Ganey, Ho Yi Wan, Samuel A. Cushman, Christina D. Vojta. 2017. Conflicting perspectives on spotted owls, wildfire, and forest restoration
• Samuel A. Cushman, Tzeidle N. Wasserman. 2017. Quantifying loss and degradation of former American marten habitat due to the impacts of forestry operations and associated road networks in northern Idaho, USA [Chapter 12]
• A. J. Shirk, E. L. Landguth, Samuel A. Cushman. 2017. A comparison of individual-based genetic distance metrics for landscape genetics
• Andrew J. Shirk, Erin L. Landguth, Samuel A. Cushman. 2017. A comparison of regression methods for model selection in individual-based landscape genetic analysis
• Helen M. Bothwell, Samuel A. Cushman, Scott A. Woolbright, Erika I. Hersch-Green, Luke M. Evans, Thomas G. Whitham, Gerard J. Allan. 2017. Conserving threatened riparian ecosystems in the American West: Precipitation gradients and river networks drive genetic connectivity and diversity in a foundation riparian tree (Populus angustifolia)
• Andrew M. Kittle, Anjali C. Watson, Samuel A. Cushman, David. W. Macdonald. 2017. Forest cover and level of protection influence the island-wide distribution of an apex carnivore and umbrella species, the Sri Lankan leopard (Panthera pardus kotiya)
• Ho Yi Wan, Kevin McGarigal, Joseph L. Ganey, Valentin Lauret, Brad C. Timm, Samuel A. Cushman. 2017. Meta-replication reveals nonstationarity in multi-scale habitat selection of Mexican Spotted Owl
• Lacy Robinson, Samuel A. Cushman, Michael K. Lucid. 2017. Winter bait stations as a multispecies survey tool
• Katherine Zeller, Kevin McGarigal, Samuel A. Cushman, Paul Beier, T. Winston Vickers, Walter M. Boyce. 2017. Sensitivity of resource selection and connectivity models to landscape definition
• Samuel A. Cushman, Ewan A. Macdonald, Erin L. Landguth, Yadvinder Malhi, David W. Macdonald. 2017. Multiple-scale prediction of forest loss risk across Borneo
• Erin L. Landguth, Zachary A. Holden, Mary F. Mahalovich, Samuel A. Cushman. 2017. Using landscape genetics simulations for planting blister rust resistant whitebark pine in the US northern Rocky Mountains
• J.-P. Puyravaud, Samuel A. Cushman, P. Davidar, D. Madappa. 2016. Predicting landscape connectivity for the Asian elephant in its largest remaining subpopulation
• Samuel A. Cushman, Erin L. Landguth. 2016. Spatially heterogeneous environmental selection strengthens evolution of reproductively isolated populations in a Dobzhansky-Muller system of hybrid incompatibility
• Kang Huang, Songtao Guo, Samuel A. Cushman, Derek W. Dunn, Xiaoguang Qi, Rong Hou, Jing Zhang, Qi Li, Qiang Zhang, Zhen Shi, Kan Zhang, Baoguo Li. 2016. Population structure of the golden snub-nosed monkey Rhinopithecus roxellana in the Qinling Mountains, central China
• Tania Barros, Samuel A. Cushman, Joao Carvalho, Carlos Fonseca. 2016. Mediterranean scrubland and elevation drive gene flow of a Mediterranean carnivore, the Egyptian mongoose Herpestes ichneumon (Herpestidae)
• E. M. Moqanaki, Samuel A. Cushman. 2016. All roads lead to Iran: Predicting landscape connectivity of the last stronghold for the critically endangered Asiatic cheetah
• Philip Riordan, Samuel A. Cushman, David Mallon, Kun Shi, Joelene Hughes. 2016. Predicting global population connectivity and targeting conservation action for snow leopard across its range
• Susana Rostro-Garcia, Lhendup Tharchen, Leandro Abade, Christos Astaras, Samuel A. Cushman, David W. Macdonald. 2016. Scale dependence of felid predation risk: Identifying predictors of livestock kills by tiger and leopard in Bhutan
• Maria Vergara, Samuel A. Cushman, Fermin Urra, Aritz Ruiz-Gonzalez. 2016. Shaken but not stirred: Multiscale habitat suitability modeling of sympatric marten species (Martes martes and Martes foina) in the northern Iberian Peninsula
• Carol L. Chambers, Samuel A. Cushman, Arnulfo Medina-Fitoria, Jose Martinez-Fonseca, Marlon Chavez-Velasquez. 2016. Influences of scale on bat habitat relationships in a forested landscape in Nicaragua
• Samuel A. Cushman. 2016. Calculating the configurational entropy of a landscape mosaic
• Kevin McGarigal, Ho Yi Wan, Katherine Zeller, Brad C. Timm, Samuel A. Cushman. 2016. Multi-scale habitat selection modeling: A review and outlook
• Niko Balkenhol, Samuel A. Cushman, Lisette P. Waits, Andrew Storfer. 2016. Current status, future opportunities, and remaining challenges in landscape genetics [Chapter 14]
• Lisette P. Waits, Samuel A. Cushman, Steve F. Spear. 2016. Applications of landscape genetics to connectivity research in terrestrial animals [Chapter 12]
• Brad C. Timm, Kevin McGarigal, Samuel A. Cushman, Joseph L. Ganey. 2016. Multi-scale Mexican spotted owl (Strix occidentalis lucida) nest/roost habitat selection in Arizona and a comparison with single-scale modeling results
• Zachary A. Holden, Alan Swanson, Anna E. Klene, John T. Abatzoglou, Solomon Z. Dobrowski, Samuel A. Cushman, John R. Squires, Gretchen Moisen, Jared W. Oyler. 2016. Development of high-resolution (250 m) historical daily gridded air temperature data using reanalysis and distributed sensor networks for the US northern Rocky Mountains
• Katherine Zeller, Kevin McGarigal, Samuel A. Cushman, Paul Beier, T. Winston Vickers, Walter M. Boyce. 2015. Using step and path selection functions for estimating resistance to movement: Pumas as a case study
• Maria C. Mateo-Sanchez, Niko Balkenhol, Samuel A. Cushman, Trinidad Perez, Ana Dominguez, Santiago Saura. 2015. A comparative framework to infer landscape effects on population genetic structure: Are habitat suitability models effective in explaining gene flow?
• Leslie C. Parks, David O. Wallin, Samuel A. Cushman, Brad H. McRae. 2015. Landscape-level analysis of mountain goat population connectivity in Washington and southern British Columbia
• Aritz Ruiz-Gonzalez, Samuel A. Cushman, Maria Jose Madeira, Ettore Randi, Benjamin J. Gomez-Moliner. 2015. Isolation by distance, resistance and/or clusters? Lessons learned from a forest-dwelling carnivore inhabiting a heterogeneous landscape
• Jiuyan Yang, Samuel A. Cushman, Xuemei Song, Jie Yang, Pujin Zhang. 2015. Genetic diversity and drivers of genetic differentiation of Reaumuria soongorica of the Inner Mongolia plateau in China
• Erin L. Landguth, Norman A. Johnson, Samuel A. Cushman. 2015. Clusters of incompatible genotypes evolve with limited dispersal
• Maria C. Mateo-Sanchez, Niko Balkenhol, Samuel A. Cushman, Trinidad Perez, Ana Dominguez, Santiago Saura. 2015. Estimating effective landscape distances and movement corridors: Comparison of habitat and genetic data
• Samuel A. Cushman. 2015. Pushing the envelope in genetic analysis of species invasion
• Samuel A. Cushman, Nicholas B. Elliot, David W. Macdonald, Andrew J. Loveridge. 2015. A multi-scale assessment of population connectivity in African lions (Panthera leo) in response to landscape change
• Andrew J. Shirk, Michael A. Schroeder, Leslie A. Robb, Samuel A. Cushman. 2015. Empirical validation of landscape resistance models: insights from the Greater Sage-Grouse (Centrocercus urophasianus)
• Samuel A. Cushman. 2015. Thermodynamics in landscape ecology: The importance of integrating measurement and modeling of landscape entropy
• Samuel A. Cushman, Jesse S. Lewis, Erin L. Landguth. 2014. Why did the bear cross the road? Comparing the performance of multiple resistance surfaces and connectivity modeling methods
• M. C. Mateo-Sanchez, Samuel A. Cushman, S. Saura. 2014. Connecting endangered brown bear subpopulations in the Cantabrian Range (north-western Spain)
• Aritz Ruiz-Gonzalez, Mikel Gurrutxaga, Samuel A. Cushman, Maria Jose Madeira, Ettore Randi, Benjamin J. Gomez-Moliner. 2014. Landscape genetics for the empirical assessment of resistance surfaces: The European pine marten (Martes martes) as a target-species of a regional ecological network
• Nicholas B. Elliot, Samuel A. Cushman, Andrew J. Loveridge, Godfrey Mtare, David W. Macdonald. 2014. Movements vary according to dispersal stage, group size, and rainfall: The case of the African lion
• Nicholas B. Elliot, Samuel A. Cushman, David W. Macdonald, Andrew J. Loveridge. 2014. The devil is in the dispersers: Predictions of landscape connectivity change with demography
• Andrew J. Shirk, Samuel A. Cushman. 2014. Spatially-explicit estimation of Wright's neighborhood size in continuous populations
• Samuel A. Cushman, Tamara Max, Nashelly Meneses, Luke M. Evans, Sharon Ferrier, Barbara Honchak, Thomas G. Whitham, Gerard J. Allan. 2014. Landscape genetic connectivity in a riparian foundation tree is jointly driven by climatic gradients and river networks
• Jessica A. Castillo, Clinton W. Epps, Anne R. Davis, Samuel A. Cushman. 2014. Landscape effects on gene flow for a climate-sensitive montane species, the American pika
• Katherine Zeller, Kevin McGarigal, Paul Beier, Samuel A. Cushman, T. Winston Vickers, Walter M. Boyce. 2014. Sensitivity of landscape resistance estimates based on point selection functions to scale and behavioral state: Pumas as a case study
• Brian K. Hand, Samuel A. Cushman, Erin L. Landguth, John Lucotch. 2014. Assessing multi-taxa sensitivity to the human footprint, habitat fragmentation and loss by exploring alternative scenarios of dispersal ability and population size: A simulation approach
• Andrew J. Shirk, Martin G. Raphael, Samuel A. Cushman. 2014. Spatiotemporal variation in resource selection: Insights from the American marten (Martes Americana)
• Samuel A. Cushman. 2014. Grand challenges in evolutionary and population genetics: The importance of integrating epigenetics, genomics, modeling, and experimentation
• Lucretia E. Olson, Joel D. Sauder, Nathan M. Albrecht, Ray S. Vinkey, Samuel A. Cushman, Michael K. Schwartz. 2014. Modeling the effects of dispersal and patch size on predicted fisher (Pekania [Martes] pennanti) distribution in the U.S. Rocky Mountains
• Samuel A. Cushman, Jesse S. Lewis, Erin L. Landguth. 2013. Evaluating the intersection of a regional wildlife connectivity network with highways
• Samuel A. Cushman, Timothy J. Mersmann, Gretchen Moisen, Kevin S. McKelvey, Christina D. Vojta. 2013. Chapter 5. Using Habitat Models for Habitat Mapping and Monitoring
• Samuel A. Cushman, Kevin McGarigal, Kevin S. McKelvey, Christina D. Vojta, Claudia M. Regan. 2013. Chapter 6. Landscape Analysis for Habitat Monitoring
• Barbara S. McCall, Michael S. Mitchell, Michael K. Schwartz, Jim Hayden, Samuel A. Cushman, Pete Zager, Wayne F. Kasworm. 2013. Combined use of mark-recapture and genetic analyses reveals response of a black bear population to changes in food productivity
• Jiuyan Yang, Samuel A. Cushman, Jie Yang, Mingbo Yang, Tiejun Bao. 2013. Effects of climatic gradients on genetic differentiation of Caragana on the Ordos Plateau, China
• Samuel A. Cushman, Erin L. Landguth, Curtis H. Flather. 2013. Evaluating population connectivity for species of conservation concern in the American Great Plains
• Kevin S. McKelvey, Jennifer E. Ramirez, Kristine L. Pilgrim, Samuel A. Cushman, Michael K. Schwartz. 2013. Genetic sampling of Palmer's chipmunks in the Spring Mountains, Nevada
• Michael Lucid, L. Robinson, Samuel A. Cushman, L. Allen, Michael K. Schwartz, Kristine L. Pilgrim. 2013. Status of fisher in the northern Idaho panhandle and adjacent mountain ranges
• Maria C. Mateo Sanchez, Samuel A. Cushman, Santiago Saura. 2013. Scale dependence in habitat selection: The case of the endangered brown bear (Ursus arctos) in the Cantabrian Range (NW Spain)
• Dawn M. Reding, Samuel A. Cushman, Todd E. Gosselink, William R. Clark. 2013. Linking movement behavior and fine-scale genetic structure to model landscape connectivity for bobcats (Lynx rufus)
• Samuel A. Cushman, Tzeidle N. Wasserman, Erin L. Landguth, Andrew J. Shirk. 2013. Re-evaluating causal modeling with mantel tests in landscape genetics
• Samuel A. Cushman, Brad McRae, Frank Adriaensen, Paul Beier, Mark Shirley, Katherine Zeller. 2013. Biological corridors and connectivity [Chapter 21]
• Samuel A. Cushman, Andrew J. Shirk, Erin L. Landguth. 2013. Landscape genetics and limiting factors
• Tzeidle N. Wasserman, Samuel A. Cushman, Jeremy S. Littell, Andrew J. Shirk, Erin L. Landguth. 2013. Population connectivity and genetic diversity of American marten (Martes americana) in the United States northern Rocky Mountains in a climate change context
• Andrew J. Shirk, Tzeidle N. Wasserman, Samuel A. Cushman, Martin G. Raphael. 2012. Scale dependency of American marten (Martes americana) habitat relations [Chapter 12]
• Deborah A. Rudnick, Sadie J. Ryan, Paul Beier, Samuel A. Cushman, Fred Dieffenbach, Clinton W. Epps, Leah R. Gerber, Joel Hartter, Jeff S. Jenness, Julia Kintsch, Adina M. Merenlender, Ryan M. Perkl, Damian V. Preziosi, Stephen C. Trombulak. 2012. The role of landscape connectivity in planning and implementing conservation and restoration priorities
• Christopher Blair, Dana E. Weigel, Matthew Balazik, Annika T. H. Keeley, Faith M. Walker, Erin Landguth, Samuel A. Cushman, Melanie Murphy, Lisette Waits, Niko Balkenhol. 2012. A simulation-based evaluation of methods for inferring linear barriers to gene flow
• Samuel A. Cushman, Erin L. Landguth. 2012. Multi-taxa population connectivity in the northern Rocky Mountains
• Samuel A. Cushman, Erin L. Landguth, Curtis H. Flather. 2012. Evaluating the sufficiency of protected lands for maintaining wildlife population connectivity in the northern Rocky Mountains
• T. N. Wasserman, Samuel A. Cushman, A. S. Shirk, E. L. Landguth, J. S. Littell. 2012. Simulating the effects of climate change on population connectivity of American marten (Martes americana) in the northern Rocky Mountains, USA
• Samuel A. Cushman, E. L. Landguth, A. J. Shirk. 2012. Simulating pattern-process relationships to validate landscape genetic models
• B. K. Hand, J. Glassy, Samuel A. Cushman, E. L. Landguth, M. A. Sawaya. 2012. UNICOR: a species connectivity and corridor network simulator
• E. L. Landguth, Samuel A. Cushman, N. Johnson. 2012. Simulating natural selection in landscape genetics
• Erin L. Landguth, Bradley C. Fedy, Sara J. Oyler-McCance, Andrew L. Garey, Sarah L. Emel, Matthew Mumma, Helene H. Wagner, Marie-Josee Fortin, Samuel A. Cushman. 2012. Effects of sample size, number of markers, and allelic richness on the detection of spatial genetic pattern
• Tabitha A. Graves, Tzeidle N. Wasserman, Milton Cezar Ribeiro, Erin L. Landguth, Stephen F. Spear, Niko Balkenhol, Colleen B. Higgins, Marie-Josee Fortin, Samuel A. Cushman, Lisette P. Waits. 2012. The influence of landscape characteristics and home-range size on the quantification of landscape-genetics relationships
• Samuel A. Cushman, Andrew J. Shirk, Erin L. Landguth. 2012. Separating the effects of habitat area, fragmentation and matrix resistance on genetic differentiation in complex landscapes
• Tzeidle N. Wasserman, Samuel A. Cushman, David O. Wallin, Jim Hayden. 2012. Multi scale habitat relationships of Martes americana in northern Idaho, U.S.A.
• Samuel A. Cushman, Erin L. Landguth. 2012. Ecological associations, dispersal ability, and landscape connectivity in the northern Rocky Mountains
• E. L. Landguth, B. K. Hand, J. M. Glassy, Samuel A. Cushman, M. Jacobi, T. J. Julian. 2011. CDPOP Users Manual
• E. L. Landguth, B. K. Hand, J. M. Glassy, Samuel A. Cushman, R. T. Carlson. 2011. UNICOR Users Manual
• Samuel A. Cushman, Martin G. Raphael, Len Ruggiero, A. S. Shirk, T. N. Wasserman, E. C. O'Doherty. 2011. Limiting factors and landscape connectivity: the American marten in the Rocky Mountains
• Samuel A. Cushman, R. Mace, T. Chilton, K. C. Kendall, E. L. Landguth, R. A. Short Bull, Michael K. Schwartz, Kevin S. McKelvey, Fred W. Allendorf, G. Luikart. 2011. Why replication is important in landscape genetics: American black bear in the Rocky Mountains
• A. J. Shirk, Samuel A. Cushman. 2011. sGD software for estimating spatially explicit indices of genetic diversity
• Jeremy S. Littell, Donald McKenzie, Becky K. Kerns, Samuel A. Cushman, Charles Shaw. 2011. Managing uncertainty in climate-driven ecological models to inform adaptation to climate change
• Jeffrey Evans, Melanie A. Murphy, Zachary A. Holden, Samuel A. Cushman. 2011. Modeling species distribution and change using random forest [Chapter 8]
• Samuel A. Cushman, Tzeidle N. Wasserman, Kevin McGarigal. 2011. Landscape fire and wildlife habitat [chapter 9]
• Samuel A. Cushman, Falk Huettmann. 2010. Future and outlook: Where are we, and where will the spatial information management in wildlife ecology be in 50 years from now? [Chapter 24]
• Zachery A. Holden, Michael A. Crimmins, Samuel A. Cushman, Jeremy S. Littell. 2010. Empirical modeling of spatial and temporal variation in warm season nocturnal air temperatures in two North Idaho mountain ranges, USA
• Samuel A. Cushman, Erin L. Landguth, Curtis H. Flather. 2010. Phase I: Climate change and connectivity: Assessing landscape and species vulnerability
• Samuel A. Cushman. 2010. Space and time in ecology: Noise or fundamental driver? [chapter 2]
• Samuel A. Cushman, Falk Huettmann. 2010. Introduction: Ecological knowledge, theory and information in space and time [Chapter 1]
• Hawthorne L. Beyer, Jeff Jenness, Samuel A. Cushman. 2010. Components of spatial information management in wildlife ecology: Software for statistical and modeling analysis [Chapter 14]
• Samuel A. Cushman, Bradley W. Compton, Kevin McGarigal. 2010. Habitat fragmentation effects depend on complex interactions between population size and dispersal ability: Modeling influences of roads, agriculture and residential development across a range of life-history characteristics [chapter 20]
• Samuel A. Cushman, Michael Chase, Curtice Griffin. 2010. Mapping landscape resistance to identify corridors and barriers for elephant movement in southern Africa [Chapter 19]
• Samuel A. Cushman. 2010. Animal movement data: GPS telemetry, autocorrelation and the need for path-level analysis [chapter 7]
• Samuel A. Cushman, Kevin S. McKelvey. 2010. Data on distribution and abundance: Monitoring for research and management [Chapter 6]
• Samuel A. Cushman, Kevin Gutzweiler, Jeffrey S. Evans, Kevin McGarigal. 2010. The Gradient Paradigm: A conceptual and analytical framework for landscape ecology [Chapter 5]
• Samuel A. Cushman, Jeffrey S. Evans, Kevin McGarigal. 2010. Landscape ecology: Past, present, and future [Chapter 4]
• Jeremy Littell, Kevin McGarigal, Samuel A. Cushman. 2010. The problem of ecological scaling in spatially complex, nonequilibrium ecological systems [chapter 3]
• Troy M. Hegel, Samuel A. Cushman, Jeffrey Evans, Falk Huettmann. 2010. Current state of the art for statistical modeling of species distributions [Chapter 16]
• Samuel A. Cushman, Jesse S. Lewis. 2010. Movement behavior explains genetic differentiation in American black bears
• Samuel A. Cushman, Jeffrey S. Evans, Kevin McGarigal, Joseph M. Kiesecker. 2010. Toward Gleasonian landscape ecology: From communities to species, from patches to pixels
• Bryan K. Epperson, Brad H. McRae, Kim Scribner, Samuel A. Cushman, Michael S. Rosenberg, Marie-Josee Fortin, Patrick M. A. James, Melanie Murphy, Stephanie Manel, Pierre Legendre, Mark R. T. Dale. 2010. Utility of computer simulations in landscape genetics
• Samuel A. Cushman, Erin L. Landguth. 2010. Scale dependent inference in landscape genetics
• Samuel A. Cushman, Erin L. Landguth. 2010. Spurious correlations and inference in landscape genetics
• Tzeidle N. Wasserman, Samuel A. Cushman, Michael K. Schwartz, David O. Wallin. 2010. Spatial scaling and multi-model inference in landscape genetics: Martes americana in northern Idaho
• A. J. Shirk, D. O. Wallin, Samuel A. Cushman, C. G. Rice, K. I. Warheit. 2010. Inferring landscape effects on gene flow: A new model selection framework
• Gernot Segelbacher, Samuel A. Cushman, Bryan K. Epperson, Marie-Josee Fortin, Olivier Francois, Olivier J. Hardy, Rolf Holderegger, Stephanie Manel. 2010. Applications of landscape genetics in conservation biology: concepts and challenges
• Erin L. Landguth, Samuel A. Cushman. 2010. CDPOP: A spatially explicit cost distance population genetics program
• E. L. Landguth, Samuel A. Cushman, M. A. Murphy, G. Luikart. 2010. Relationships between migration rates and landscape resistance assessed using individual-based simulations
• Michael K. Schwartz, Gordon Luikart, Kevin S. McKelvey, Samuel A. Cushman. 2010. Landscape genomics: A brief perspective [Chapter 9]
• E. L. Landguth, Samuel A. Cushman, Michael K. Schwartz, Kevin S. McKelvey, M. Murphy, G. Luikart. 2010. Quantifying the lag time to detect barriers in landscape genetics
• Samuel A. Cushman, Kevin S. McKelvey, Barry R. Noon, Kevin McGarigal. 2010. Use of abundance of one species as a surrogate for abundance of others
• Kevin S. McKelvey, Samuel A. Cushman, Michael K. Schwartz. 2009. Landscape genetics [Chapter 17]
• Jeffrey S. Evans, Samuel A. Cushman. 2009. Gradient modeling of conifer species using random forests
• Niko Balkenhol, Felix Gugerli, Samuel A. Cushman, Lisette P. Waits, Aurelie Coulon, J. W. Arntzen, Rolf Holderegger, Helene H. Wagner. 2009. Identifying future research needs in landscape genetics: Where to from here?
• Michael K. Schwartz, Jeffrey P. Copeland, Neil J. Anderson, John R. Squires, Robert M. Inman, Kevin S. McKelvey, Kristy L. Pilgrim, Lisette P. Waits, Samuel A. Cushman. 2009. Wolverine gene flow across a narrow climatic niche
• Kevin McGarigal, Sermin Tagil, Samuel A. Cushman. 2009. Surface metrics: An alternative to patch metrics for the quantification of landscape structure
• Kevin S. McKelvey, Samuel A. Cushman, Michael K. Schwartz, Leonard F. Ruggiero. 2009. Wildlife monitoring across multiple spatial scales using grid-based sampling
• Donald McKenzie, Crystal Raymond, Samuel A. Cushman. 2008. Modeling understory vegetation and its response to fire [Chapter 15]
• Samuel A. Cushman, Kevin McGarigal. 2008. Landscape metrics, scales of resolution
• Melanie A. Murphy, Jeffrey S. Evans, Samuel A. Cushman, Andrew Storfer. 2008. Representing genetic variation as continuous surfaces: An approach for identifying spatial dependency in landscape genetic studies
• Samuel A. Cushman, Kevin McGarigal, Maile C. Neel. 2008. Parsimony in landscape metrics: Strength, universality, and consistency
• Samuel A. Cushman, Kevin S. McKelvey, Michael K. Schwartz. 2008. Case study 6.1: DNA survey for fisher in northern Idaho
• Samuel A. Cushman, Kevin S. McKelvey, Michael K. Schwartz. 2008. Use of empirically derived source-destination models to map regional conservation corridors
• Kevin S. McKelvey, Curtis H. Flather, Samuel A. Cushman, Kevin McGarigal. 2008. Do forest community types provide a sufficient basis to evaluate biological diversity?
• Samuel A. Cushman, Kevin McGarigal. 2007. Multivariate landscape trajectory analysis: An example using simulation modeling of American marten habitat change under four timber harvest scenarios
• Bradley W. Compton, Kevin McGarigal, Samuel A. Cushman, Lloyd R. Gamble. 2007. A resistant-kernel model of connectivity for amphibians that breed in vernal pools
• Samuel A. Cushman, Donald McKenzie, David L. Peterson, Jeremy Littell, Kevin S. McKelvey. 2007. Research agenda for integrated landscape modeling
• Michael K. Schwartz, Samuel A. Cushman, Kevin S. McKelvey, Jim Hayden, Cory Engkjer. 2006. Detecting genotyping errors and describing black bear movement in northern Idaho
• Samuel A. Cushman, Kevin S. McKelvey. 2006. Developing statistical wildlife habitat relationships for assessing cumulative effects of fuels treatments: Final Report for Joint Fire Science Program Project
• Samuel A. Cushman, Kevin S. McKelvey, Jim Hayden, Michael K. Schwartz. 2006. Gene flow in complex landscapes: Testing multiple hypotheses with causal modeling
• Samuel A. Cushman, Donald McKenzie, David L. Peterson, Jeremy Littell, Kevin S. McKelvey. 2006. Research agenda for integrated landscape modeling
• Samuel A. Cushman. 2006. Effects of habitat loss and fragmentation on amphibians: A review and prospectus
• Kevin McGarigal, Samuel A. Cushman, Claudia Regan. 2005. Quantifying terrestrial habitat loss and fragmentation: A protocol
• Kevin McGarigal, Samuel A. Cushman. 2005. The gradient concept of landscape structure [Chapter 12]
• Samuel A. Cushman, Michael Chase, Curtice Griffin. 2005. Elephants in space and time
• Samuel A. Cushman, Kevin McGarigal. 2004. Hierarchical analysis of forest bird species-environment relationships in the Oregon Coast Range
• Samuel A. Cushman, Kevin McGarigal. 2004. Patterns in the species-environment relationship depend on both scale and choice of response variables
• Maile C. Neel, Kevin McGarigal, Samuel A. Cushman. 2004. Behavior of class-level landscape metrics across gradients of class aggregation and area
• Joanna Grand, John Buonaccorsi, Samuel A. Cushman, Curtice R. Griffin, Maile C. Neel. 2004. A multiscale landscape approach to predicting bird and moth rarity hotspots in a threatened pitch pine-scrub oak community
• Samuel A. Cushman, Kevin McGarigal. 2003. Landscape-level patterns of avian diversity in the Oregon Coast Range

Citations of Non-Forest Service Publications

• The central role of connectivity in conservation geopolitics
• Understanding climate change impacts to white pines
• Toolkit for supporting conservation decision makers in Sabah, Borneo
• Toolkit for supporting conservation decision makers in Myanmar
• Population connectivity, gene flow and climate change

Articles

• Article
  

  New Mexico's state-wide connectivity conservation initiative

  https://research.fs.usda.gov/rmrs/articles/new-mexicos-state-wide-connectivity-conservation-initiative

Multimedia

• Webinars

  Through the smoke: Spotted owls, wildfire, and forest restoration

  https://research.fs.usda.gov/rmrs/products/multimedia/webinars/through-smoke-spotted-owls-wildfire-and-forest-restoration

• University of Massachusetts, Ph.D., Landscape Ecology, 2003
• Western Washington University, Bellingham, WA, M.S., Landscape Ecology, 1997
• Evergreen State College, Olympia, WA, B.S., Landscape Ecology, 1995