Walter Shortle, PhD

Treesearch Publications

• Walter Shortle, Kevin T. Smith, Andrei G. Lapenis. 2017. Dendrochemical evidence for soil recovery from acidic deposition in forests of the northeastern U.S. with comparisons to the southeastern U.S. and Russia
• Gregory B. Lawrence, Paul W. Hazlett, Ivan J. Fernandez, Rock Ouimet, Scott Bailey, Walter Shortle, Kevin T. Smith, Michael R. Antidormi. 2015. Declining acidic deposition begins reversal of forest-soil acidification in the northeastern U.S
• Walter Shortle, Kevin T. Smith. 2015. Wood decay fungi restore essential calcium to acidic soils in northern New England
• Walter Shortle, Jody Jellison, Kevin T. Smith, Jonathan S. Schilling. 2014. Studying the role of wood-decay fungi in calcium cycling on the Penobscot Experimental Forest: a progress report
• Kevin T. Smith, Kenneth Dudzik, Walter Shortle. 2014. Tree survival 15 years after the ice storm of January 1998
• Kevin T. Smith, Jean Christophe Balouet, Walter Shortle, Michel Chalot, François Beaujard, Hakan Grudd, Don A. Vroblesky, Joel G. Burken. 2014. Dendrochemical patterns of calcium, zinc, and potassium related to internal factors detected by energy dispersive X-ray fluorescence (EDXRF)
• Kevin T. Smith, Walter Shortle. 2013. Calcium amendment may increase hydraulic efficiency and forest evapotranspiration
• Andrei G. ​Lapenis, Gregory B. Lawrence, Alexander Heim, Chengyang Zheng, Walter Shortle. 2013. Climate warming shifts carbon allocation from stemwood to roots in calcium-depleted spruce forests
• Gregory B. Lawrence, Walter Shortle, Mark B. David, Kevin T. Smith, Richard A. Warby, Andrei G. Lapenis. 2012. Early indications of soil recovery from acidic deposition in U.S. red spruce forests
• Walter Shortle, Kenneth Dudzik. 2012. Wood decay in living and dead trees: A pictorial overview
• Walter Shortle, Kevin T. Smith, Jody Jellison, Jonathan S. Schilling. 2012. Potential of decaying wood to restore root-available base cations in depleted forest soils
• Walter Shortle, Kenneth Dudzik, Kevin T. Smith. 2010. Development of wood decay in wound-initiated discolored wood of eastern red cedar
• Kevin T. Smith, Walter Shortle, Jon H. Connolly, Rakesh Minocha, Jody Jellison. 2009. Calcium fertilization increases the concentration of calcium in sapwood and calcium oxalate in foliage of red spruce
• Walter Shortle, Peter S. Murdoch, Kevin T. Smith, Rakesh Minocha, Gregory B. Lawrence. 2008. 6.0 Monitoring recovery from calcium depletion and nitrogen saturation
• Kevin T. Smith, Walter Shortle, Jody Jellison, Jon Connolly, Jonathan Schilling. 2007. Concentrations of Ca and Mg in early stages of sapwood decay in red spruce, eastern hemlock, red maple, and paper birch
• Andy B. Reinmann, Laura S. Kenefic, Ivan J. Fernandez, Walter Shortle. 2005. Soil Characteristics 30-years Post-harvest on Maine Spruce Flats: Preliminary Results
• Dirk Dujesiefken, Walter Liese, Walter Shortle, Rakesh Minocha. 2005. Response of beech and oaks to wounds made at different times of the year
• Andy Reinmann, Laura S. Kenefic, Ivan Fernandez, Walter Shortle. 2004. Preliminary Results From a Retrospective Study of Harvest Intensity, Site Productivity, and Red Spruce Growth
• Kevin T. Smith, Walter Shortle. 2003. Radial growth of hardwoods following the 1998 ice storm in New Hampshire and Maine
• Walter Shortle, Kevin T. Smith, Kenneth Dudzik. 2003. Tree Survival and Growth Following Ice Storm Injury
• Samuel B. McLaughlin, Walter Shortle, Kevin T. Smith. 2002. Dendroecological applications in air pollution and environmental chemistry: research needs
• Kevin T. Smith, Walter Shortle. 2001. Conservation of element concentration in xylem sap of red spruce
• Walter Shortle. 2000. CODIT
• T.R. Filley, P.G. Hatcher, Walter Shortle. 2000. The application of ¹³C-labeled tetramethylammonium hydroxide (¹³C-TMAH) thermochemolysis to study fungal degradation of wood
• Walter Shortle. 2000. Shigometer
• Walter Shortle, Kevin T. Smith, Rakesh Minocha, Subhash Minocha, Philip M. Wargo, Kristina A. Vogt. 2000. Tree health and physiology in a changing environment
• Walter Shortle, Rakesh Minocha. 1999. Taxol: a review
• Gregory B. Lawrence, Mark B. David, Walter Shortle, Scott Bailey, Gary M. Lovett. 1999. Mechanisms of base-cation depletion by acid deposition in forest soils of the northeastern U.S.
• Walter Shortle, Kevin T. Smith, Rakesh Minocha, Subhash C. Minocha, Philip M. Wargo, Kristiina A. Vogt. 1999. Tree health and physiology in a changing environment
• Jon H. Connolly, Walter Shortle, Jody Jellison. 1999. Translocation and incorporation of strontium carbonate derived strontium into calcium oxalate crystals by the wood decay fungus Resinicium bicolor
• N. Torelli, Walter Shortle, K. Cufar, F. Ferlin, Kevin T. Smith. 1999. Detecting changes in tree health and productivity in silver fir-beech forests of Slovenia
• Primoz Oven, Niko Torelli, Walter Shortle, Martin Zupancic. 1999. The formation of a ligno-suberised layer and necrophylactic periderm in beech bark (Fagus sylvatica L.)
• Kevin T. Smith, Walter Shortle. 1998. A first look at tree decay. An introduction to how injury and decay affect trees
• Rakesh Minocha, Walter Shortle, Gregory B. Lawrence, Mark B. David, Subhash C. Minocha. 1997. Relationships among foliar chemistry, foliar polyamines, and soil chemistry in red spruce trees growing across the northeastern United States
• Walter Shortle, Kevin T. Smith, Rakesh Minocha, Gregory B. Lawrence, Mark B. David. 1997. Acidic deposition, cation mobilization, and biochemical indicators of stress in healthy red spruce
• A. Ostrofsky, J. Jellison, Kevin T. Smith, Walter Shortle. 1997. Changes in cation concentrations in red spruce wood decayed by brown rot and white rot fungi
• Gergory B. Lawrence, Mark B. David, Scott Bailey, Walter Shortle. 1997. Assessment of calcium status in soils of red spruce forests in the northeastern United States
• Mark B. David, Gregory B. Lawrence, Walter Shortle, Scott Bailey. 1996. Calcium status of the forest floor in red spruce forests of the northeastern U.S. - past, present and future
• Gregory B. Lawrence, Mark B. David, Walter Shortle. 1996. Aluminum mobilization and calcium depletion in the forest floor of red spruce forests in the northeastern United States
• Rakesh Minocha, Walter Shortle, Gregory B. Lawrence, Mark B. David, Subhash C. Minocha. 1996. Putrescine: a marker of stress in red spruce trees
• Rakesh Minocha, Walter Shortle, Daniel J. Jr. Coughin, Subhash C. Minocha. 1996. Effects of aluminum on growth, polyamine metabolism, and inorganic ions in suspension cultures of red spruce (Picea rubens)
• Rakesh Minocha, Walter Shortle, Gregory B. Lawrence, B. David Mark, Subhash C. Minocha. 1996. Putrescine: a marker of stress in red spruce trees
• Kevin T. Smith, Walter Shortle, Rakesh Minocha, Vladislav A. Alexeyev. 1996. Dynamics of calcium concentration in stemwood of red spruce and Siberian fir
• Kevin T. Smith, Walter Shortle. 1996. Tree biology and dendrochemistry
• Walter Shortle, Kevin T. Smith, Kenneth Dudzik. 1996. Decay diseases of stemwood: Detection, diagnosis, and management
• Walter Shortle, Kevin T. Smith, Rakesh Minocha, V.A. Alexeyev. 1995. Similar patterns of change in stemwood calcium concentration in red spruce and Siberian fir
• Walter Shortle, Kevin T. Smith, Kenneth Dudzik, Sharon Parker. 1995. Response of maple sapwood to injury and infection
• Kevin T. Smith, Walter Shortle, W.D. Ostrofsky. 1995. Aluminum and calcium in fine root tips of red spruce collected from the forest floor
• Gregory B. Lawrence, Mark B. David, Walter Shortle. 1995. A new mechanism for calcium loss in forest-floor soils
• Rakesh Minocha, Walter Shortle, Subhash C. Minocha. 1994. Somatic embryogenesis in Picea abies (Norway spruce) and Picea Rubens (red spruce): growth rates and polyamines
• Rakesh Minocha, Walter Shortle. 1994. Polyamines and inorganic ions extracted from woody tissues by freeze-thawing
• Rakesh Minocha, Walter Shortle, Stephanie L. Long, Subhash C. Minocha. 1994. A rapid and reliable procedure for extraction of cellular polyamines and inorganic ions from plant tissues
• Rakesh Minocha, Walter Shortle. 1993. Fast, safe, and reliable methods for extraction of major inorganic cations from small quantities of woody plant tissues
• Rakesh Minocha, Subhash C. Minocha, Stephanie Long, Walter Shortle. 1992. Effects of aluminum on DNA synthesis, cellular polyamines, polyamine biosynthetic enzymes and inorganic ions in cell suspension cultures of a woody plant, Catharanthus roseus
• Rakesh Minocha, S.C. Minocha, A. Komamine, Walter Shortle. 1991. Regulation of DNA synthesis and cell division by polyamines in Catharanthus roseus suspension cultures
• Rakesh Minocha, Subhash C. Minocha, Atsushi Komamine, Walter Shortle. 1990. Role of polyamines in DNA synthesis of Catharanthus roseus cells grown in suspension culture
• Walter Shortle, Rakesh Minocha. 1990. Applications of ion chromatography to study pollution effects on forest trees
• Walter Shortle, Kevin T. Smith. 1988. Aluminum-induced calcium deficiency syndrome in declining red spruce
• Alex L. Shigo, Walter Shortle. 1985. Spruce Budworms Handbook: shigometry – a reference guide
• P. W. Garrett, W. K. Randall, A. L. Shigo, Walter Shortle. 1979. Inheritance of compartmentalization of wounds in sweetgum (Liquidambar styraciflua L.) and eastern cottonwood (Populus deltoides Bartr.)
• Alex L. Shigo, Walter Shortle, Julian Ochrymowych. 1977. Detection of active decay at groundline in utility poles

• North Carolina State University, Raleigh, Ph.D., Plant Pathology, 1974
• University of New Hampshire, Durham, M.S., Botany, 1970
• University of New Hampshire, Durham, B.S., Botany, 1968