Alejandro A. Royo, PhD

Treesearch Publications

• Patricia Raymond, Emilie Champagne, Daniel Dumais, Christel C. Kern, Catherine Perie, Alison D. Munson, Jean-Pierre Tremblay, Alejandro A. Royo. 2025. Moving up north: How do translocated seedlings perform in mixed-species plantings at the boreal-temperate interface?
• Andrea Brandon, Alejandro A. Royo, Samuel Reed. 2025. Restoring Future Forests: How Fire, Deer, and Canopy Gaps Shape Forest Seed Banks and Regeneration Potential
• Lance A. Vickers, Alejandro A. Royo, Susan L. Stout, Todd Ristau, William K. Oldland, Benjamin O. Knapp, Lauren S. Pile, Andrea Hille. 2025. Evaluating the extent and severity of black cherry regeneration declines in eastern forests and exploring potential causes
• Samuel P. Reed, Alejandro A. Royo, Walter P. Carson, Castilleja F. Olmsted, Lee E. Frelich, Peter B. Reich. 2025. Multiple disturbances, multiple legacies: fire, canopy gaps and deer jointly change the forest seed bank
• Laurel M. Glover, Sabrina N. Volponi, Alejandro A. Royo, Tamara L. Johnstone-Yellin. 2024. Effect of Herbivory on the Nutritional Value of Six Hardwood Species for White-Tailed Deer in the Allegheny Forest
• Bryce T. Adams, Alejandro A. Royo, Christel C. Kern, Dustin Bronson, Stephen Matthews, Andrew Gougherty, Anantha M. Prasad, Louis Iverson, Evan Rehm, Patricia Raymond, Catherine Périé, Jessica R. Miesel, Amanda M. McGraw. 2024. Identifying climatically-compatible seedlots for the eastern US: building the predictive tools and knowledge to enable forest assisted migration
• Andrew D. Bower, Katie L. Frerker, Carolyn C. Pike, Nicholas R. Labonte, Brian J. J. Palik, Alejandro A. Royo, Sarah Anderson, Arnaldo R. Ferreira, Leslie Brandt. 2024. A practical framework for applied forestry assisted migration
• Todd Hutchinson, Bryce T. Adams, Matthew B. Dickinson, Maryjane Heckel, Alejandro A. Royo, Melissa A. Thomas‐Van Gundy. 2024. Sustaining eastern oak forests: Synergistic effects of fire and topography on vegetation and fuels
• Anthony C. Yacucci, Walter P. Carson, Jenna C. Martineau, Curtis D. Burns, Brian P. Riley, Alejandro A. Royo, Thomas P. Diggins, Ian J. Renne. 2024. Native tree species prosper while exotics falter during gap-phase regeneration, but only where deer densities are near historical levels
• Baptiste Brault, Jean-Pierre Tremblay, Nelson Thiffault, Alejandro A. Royo, Steeve D. Côté. 2023. Successful forest restoration using plantation at high deer density: how neighboring vegetation drives browsing pressure and tree growth
• Kathleen Knight, Charles E. Flower, Jason S. Kilgore, Alejandro A. Royo, Timothy Fox, Rachel H. Kappler, Brian M. Hoven, Joshua Wigal, Jordan D. Bernard, Robert P. Long. 2023. Ash tree decline and mortality in Ohio and the Allegheny National Forest in Pennsylvania
• Alejandro A. Royo, Patricia Raymond, Christel C. Kern, Bryce T. Adams, Dustin Bronson, Emilie Champagne, Daniel Dumais, Eric J. Gustafson, Paula E. Marquardt, Amanda M. McGraw, Jessica R. Miesel, Alison D. Munson, Catherine Périé, Felipe J. Tavares Moreira, Anne Ola, Mathieu Bouchard, Jean-François Bissonnette. 2023. Desired REgeneration through Assisted Migration (DREAM): Implementing a research framework for climate-adaptive silviculture
• Emilie Champagne, Alejandro A. Royo, Jean-Pierre Tremblay, Patricia Raymond. 2023. Tree Assisted Migration in a Browsed Landscape: Developing an Assessment of Susceptibility
• Susan L. Stout, Andrea T. Hille, Alejandro A. Royo. 2023. Science-Management Collaboration is Essential to Address Current and Future Forestry Challenges
• Aaron Knauer, Tiffany Betras, Alejandro A. Royo, Thomas P. Diggins, Walter P. Carson. 2023. Understory plant communities fail to recover species diversity after excluding deer for nearly 20 years
• Tyler K. Refsland, Bryce T. Adams, Dustin Bronson, Christel C. Kern, Paula Marquardt, Amanda M. McGraw, Alejandro A. Royo, Jessica R. Miesel. 2023. Synthesis of plant-soil feedback effects on eastern North American tree species: implications for climate-adaptive forestry
• Michelle Elise Spicer, Alejandro A. Royo, John W. Wenzel, Walter P. Carson. 2023. Understory plant growth forms respond independently to combined natural and anthropogenic disturbances
• David W. Kramer, Thomas J. Prebyl, Nathan P. Nibbelink, Karl V Miller, Alejandro A. Royo, Jacqueline L. Frair. 2022. Managing Moose from Home: Determining Landscape Carrying Capacity for Alces alces Using Remote Sensing
• Jennifer M. Nieves, Jeffrey S. Ward, Alejandro A. Royo, Marc E. McDill, Jesse K. Kreye, Kim C. Steiner. 2022. Stand and site characteristics affect the probability of stump sprouting in some eastern North American hardwoods
• Alejandro A. Royo, Walter P. Carson. 2022. Stasis in forest regeneration following deer exclusion and understory gap creation: A 10‐year experiment
• Cornelia Wilson, Alejandro A. Royo, John Stanovick, Scott E. Schlarbaum, Ami M. Sharp, Sandra L. Anagnostakis. 2022. Deer browse susceptibility limits chestnut restoration success in northern hardwood forests
• Samuel P. Reed, Alejandro A. Royo, Alexander T. Fotis, Kathleen Knight, Charles E. Flower, Peter S. Curtis. 2022. The long‐term impacts of deer herbivory in determining temperate forest stand and canopy structural complexity
• Emilie Champagne, Patricia Raymond, Alejandro A. Royo, James D. M. Speed, Jean-Pierre Tremblay, Steeve D. Côté. 2021. A Review of Ungulate Impacts on the Success of Climate-Adapted Forest Management Strategies
• Emilie Champagne, Alejandro A. Royo, Jean-Pierre Tremblay, Patricia Raymond. 2021. Tree assisted migration in a browsed landscape: Can we predict susceptibility to herbivores?
• Alejandro A. Royo, Lance Vickers, Robert P. Long, Todd Ristau, Scott H. Stoleson, Susan L. Stout. 2021. The Forest of Unintended Consequences: Anthropogenic Actions Trigger the Rise and Fall of Black Cherry
• Laurent De Vriendt, Nelson Thiffault, Alejandro A. Royo, Martin Barrette, Jean-Pierre Tremblay. 2020. Moose Browsing Tends Spruce Plantations More Efficiently Than a Single Mechanical Release
• Cornelia Wilson, Alejandro A. Royo, Scott E. Schlarbaum, Matthew Peters, Ami M. Sharp, Sandra L. Anagnostakis. 2020. The effect of site quality on performance of American Chestnut (Castanea dentata) seedlings bred for blight (Cryphonectria parasitica) resistance
• Todd Ristau, Alejandro A. Royo. 2020. Influence of stand age, soil attributes, and cover type on Rubus (Rosaceae) seed bank abundance
• Emilie Champagne, Michaël Bonin, Alejandro A. Royo, Jean-Pierre Tremblay, Patricia Raymond. 2020. Predicting terpene content in dried conifer shoots using near infrared spectroscopy
• Emilie Champagne, Alejandro A. Royo, Jean-Pierre Tremblay, Patricia Raymond. 2019. Phytochemicals Involved in Plant Resistance to Leporids and Cervids: a Systematic Review
• Alejandro A. Royo, Susan L. Stout. 2019. Recognition, response, and recovery: Deer impact research in Allegheny hardwood forests
• Susan L. Stout, Patrick Brose, Helene Cleveland, Robert P. Long, Barbara McGuinness, Matthew Peters, Joanne Rebbeck, Todd Ristau, Alejandro A. Royo, Scott H. Stoleson, Scott Thomasma, Mark Twery, Sarah Wurzbacher. 2019. Fifty years of science-management cooperation from the SILVAH community of practice
• Jacob B. Slyder, John W. Wenzel, Alejandro A. Royo, Michelle Elise Spicer, Walter P. Carson. 2019. Post-windthrow salvage logging increases seedling and understory diversity with little impact on composition immediately after logging
• Lance Vickers, William McWilliams, Benjamin Knapp, Anthony D’Amato, Daniel Dey, Yvette Dickinson, John M. Kabrick, Laura S. Kenefic, Christel C. Kern, David Larsen, Alejandro A. Royo, Michael Saunders, Stephen Shifley, James A. Westfall. 2019. Are Current Seedling Demographics Poised to Regenerate Northern US Forests?
• Alejandro A. Royo, John Stanovick. 2019. Deer browsing overwhelms extended leaf phenology benefits: A test case with Rubus allegheniensis and a recalcitrant hay-scented fern layer
• Alejandro A. Royo, Cornelia Wilson, John Stanovick, Susan L. Stout. 2019. Timing is Not Everything: Assessing the Efficacy of Pre- Versus Post-Harvest Herbicide Applications in Mitigating the Burgeoning Birch Phenomenon in Regenerating Hardwood Stands
• Quinn Morgan, Tamara L. Johnstone-Yellin, Cornelia Wilson, Matthew Peters, Alejandro A. Royo. 2019. Partitioning and predicting forage biomass from total aboveground biomass of regenerating tree species using dimensional analyses
• William McWilliams, Jim Westfall, Patrick Brose, Daniel C. C. Dey, Anthony W. D'Amato, Yvette L. Dickinson, Mary Ann Fajvan, Laura S. Kenefic, Christel C. Kern, Kenneth M. Laustsen, Shawn L. Lehman, Randall Morin, Todd Ristau, Alejandro A. Royo, Andrew M. Stoltman, Susan L. Stout. 2018. Subcontinental-scale patterns of large-ungulate herbivory and synoptic review of restoration management implications for midwestern and northeastern forests
• Patricia Raymond, Alejandro A. Royo, Marcel Prévost, Daniel Dumais. 2018. Assessing the single-tree and small group selection cutting system as intermediate disturbance to promote regeneration and diversity in temperate mixedwood stands
• Alexandro B. Leverkus, José María Rey Benayas, Jorge Castro, Dominique Boucher, Stephen Brewer, Brandon M. Collins, Daniel Donato, Shawn Fraver, Barbara E. Kishchuk, Eun-Jae Lee, David B. Lindenmayer, Emanuele Lingua, Ellen Macdonald, Raffaella Marzano, Charles C. Rhoades, Alejandro A. Royo, Simon Thorn, Joseph Wagenbrenner, Kaysandra Waldron, Thomas Wohlgemuth, Lena Gustafsson. 2018. Salvage logging effects on regulating and supporting ecosystem services — a systematic map
• Charles E. Flower, Jeremie Fant, Sean Hoban, Kathleen Knight, Laura Steger, Elijah Aubihl, Miquel Gonzalez-Meler, Stephen Forry, Andrea Hille, Alejandro A. Royo. 2018. Optimizing conservation strategies for a threatened tree species: In situ conservation of white ash (Fraxinus americana L.) genetic diversity through insecticide treatment
• Kristine M Averill, David A Mortensen, Erica A H Smithwick, Susan Kalisz, William J McShea, Norman A Bourg, John D Parker, Alejandro A. Royo, Marc D Abrams, David K Apsley, Bernd Blossey, Douglas H Boucher, Kai L Caraher, Antonio DiTommaso, Sarah E Johnson, Robert Masson, Victoria A. Nuzzo. 2017. A regional assessment of white-tailed deer effects on plant invasion
• Alejandro A. Royo, David W. Kramer, Karl V. Miller, Nathan P. Nibbelink, Susan L. Stout. 2017. Spatio-temporal variation in foodscapes modifies deer browsing impact on vegetation
• Katherine Sinacore, Jefferson Scott Hall, Catherine Potvin, Alejandro A. Royo, Mark J. Ducey, Mark S. Ashton, Shijo Joseph. 2017. Unearthing the hidden world of roots: Root biomass and architecture differ among species within the same guild
• Charles E. Flower, Elijah Aubihl, Jeremie Fant, Stephen Forry, Andrea Hille, Kathleen Knight, William K. Oldland, Alejandro A. Royo, Richard M. Turcotte. 2017. In-situ genetic conservation of white ash (Fraxinus americana) at the Allegheny national forest
• Martin Barrette, Louis Bélanger, Louis De Grandpré, Alejandro A. Royo. 2017. Demographic disequilibrium caused by canopy gap expansion and recruitment failure triggers forest cover loss
• William McWilliams, Jim Westfall, Patrick Brose, Shawn L. Lehman, Randall Morin, Todd Ristau, Alejandro A. Royo, Susan L. Stout. 2017. After 25 years, what does the Pennsylvania Regeneration Study tell us about oak/hickory forests under stress?
• Nathan R. De Jager, Patrick J. Drohan, Brian R. Miranda, Brian R. R. Sturtevant, Susan L. Stout, Alejandro A. Royo, Eric J. Gustafson, Mark C. Romanski. 2017. Simulating ungulate herbivory across forest landscapes: A browsing extension for LANDIS-II
• Christel C. Kern, Julia I. Burton, Patricia Raymond, Anthony W. D'Amato, William S. Keeton, Alejandro A. Royo, Michael B. Walters, Christopher R. Webster, John L. Willis. 2017. Challenges facing gap-based silviculture and possible solutions for mesic northern forests in North America
• Thomas H. Pendergast, Shane M. Hanlon, Zachary M. Long, Alejandro A. Royo, Walter P. Carson. 2016. The legacy of deer overabundance: long-term delays in herbaceous understory recovery
• Alejandro A. Royo, Chris J. Peterson, John Stanovick, Walter P. Carson. 2016. Evaluating the ecological impacts of salvage logging: can natural and anthropogenic disturbances promote coexistence?
• Alejandro A. Royo, David W. Kramer, Karl V. Miller, Nathan P. Nibbelink, Susan L. Stout. 2016. The canary in the coal mine: Sprouts as a rapid indicator of browse impact in managed forests
• Michael J. Chips, Ellen H. Yerger, Arpad Hervanek, Tim Nuttle, Alejandro A. Royo, Jonathan N. Pruitt, Terrence P. McGlynn, Cynthia L. Riggall, Walter P. Carson. 2015. The indirect impact of long-term overbrowsing on insects in the Allegheny National Forest region of Pennsylvania
• William McWilliams, Jim Westfall, Patrick Brose, Daniel C. C. Dey, Mark A. Hatfield, Katherine Johnson, Kenneth M. Laustsen, Shawn L. Lehman, Randall Morin, Mark D. D. Nelson, Todd Ristau, Alejandro A. Royo, Susan L. Stout, Tom W. Willard, Christopher W. W. Woodall. 2015. A regeneration indicator for Forest Inventory and Analysis: history, sampling, estimation, analytics, and potential use in the midwest and northeast United States
• Cynthia Huebner, Jim Steinman, Todd Hutchinson, Todd Ristau, Alejandro A. Royo. 2014. The distribution of a non-native (Rosa multiflora) and native (Kalmia latifolia) shrub in mature closed-canopy forests across soil fertility gradients
• Tim Nuttle, Todd Ristau, Alejandro A. Royo. 2014. Long-term biological legacies of herbivore density in a landscape-scale experiment: forest understories reflect past deer density treatments for at least 20 years
• Susan L. Stout, Alejandro A. Royo, David S. deCalesta, Kevin McAleese, James C. Finley. 2013. The Kinzua Quality Deer Cooperative: can adaptive management and local stakeholder engagement sustain reduced impact of ungulate browsers in forest systems?
• Chris J. Peterson, Lisa M. Krueger, Alejandro A. Royo, Scott Stark, Walter P. Carson. 2013. Disturbance size and severity covary in small and mid-size wind disturbances in Pennsylvania northern hardwoods forests
• Tim Nuttle, Alejandro A. Royo, Mary Beth Adams, Walter P. Carson. 2013. Historic disturbance regimes promote tree diversity only under low browsing regimes in eastern deciduous forest
• Alejandro A. Royo, Todd Ristau. 2013. Stochastic and deterministic processes regulate spatio-temporal variation in seed bank diversity
• Cynthia Huebner, Todd Hutchinson, Todd Ristau, Alejandro A. Royo, James Steinman. 2012. Documenting the Regional and local distribution of Kalmia latifolia and Rosa multiflora in West Virginia, Ohio, and Pennsylvania Forests along a soil fertility gradient
• Alejandro A. Royo, Kathleen Knight. 2012. White ash (Fraxinus americana) decline and mortality: the role of site nutrition and stress history
• Daniel C. C. Dey, Alejandro A. Royo, Emile S. Gardiner, Luis Mario Cardenas. 2012. Sustaining Quercus humboldtii and Colombobalanus excelsa on the Colombian landscape: preservation or conservation - a research perspective
• Alejandro A. Royo, Kathleen Knight, Jamie M. Himes, Ashley N. Will. 2012. White ash (Fraxinus americana) health in the Allegheny plateau region, Pennsylvania: Evaluating the relationship between FIA phase 3 crown variables and a categorical rating system
• Daniel C. C. Dey, Alejandro A. Royo, Emile S. Gardiner, Luis Mario Cardenas. 2012. Sustaining Quercus humboltii and Colombobalanus excelsa on the Colombian landscape: preservation or conservation - a research perspective
• Morgan Kain, Loretta Battaglia, Alejandro A. Royo, Walter P. Carson. 2011. Over-browsing in Pennsylvania creates a depauperate forest dominated by an understory tree: results from a 60-year-old deer exclosure
• Chandra Goetsch, Jennifer Wigg, Alejandro A. Royo, Todd Ristau, Walter P. Carson. 2011. Chronic over browsing and biodiversity collapse in a forest understory in Pennsylvania: Results from a 60 year-old deer exclusion plot
• Alejandro A. Royo, Tamara Heartsill Scalley, Samuel Moya, Fred N. Scatena. 2011. Non-arborescent vegetation trajectories following repeated hurricane disturbance: ephemeral versus enduring responses
• Daniel C. C. Dey, Alejandro A. Royo, Patrick Brose, Todd Hutchinson, Marty Spetich, Scott H. Stoleson. 2010. An ecologically based approach to oak silviculture: a synthesis of 50 years of oak ecosystem research in North America
• Kathleen Knight, Daniel A. Herms, John Cardina, Robert P. Long, Joanne Rebbeck, Kamal J.K. Gandhi, Annemarie Smith, Wendy S. Klooster, Catherine P. Herms, Alejandro A. Royo. 2010. Emerald ash borer aftermath forests: The dynamics of ash mortality and the responses of other plant species
• Kurt O. Reinhart, Alejandro A. Royo, Stacie A. Kageyama, Keith Clay. 2010. Canopy gaps decrease microbial densities and disease risk for a shade-intolerant tree species
• Alejandro A. Royo, Walter P. Carson. 2010. The formation of dense understory layers in forests worldwide: consequences and implications for forest dynamics, biodiversity, and succession
• Alejandro A. Royo, Susan L. Stout, David S. deCalesta, Timothy G. Pierson. 2010. Restoring forest herb communities through landscape-level deer herd reductions: Is recovery limited by legacy effects?
• Alejandro A. Royo, Rachel Collins, Mary Beth Adams, Chad Kirschbaum, Walter P. Carson. 2010. Pervasive interactions between ungulate browsers and disturbance regimes promote temperate forest herbaceous diversity
• Lisa M. Krueger, Chris J. Peterson, Alejandro A. Royo, Walter P. Carson. 2009. Evaluating relationships among tree growth rate, shade tolerance, and browse tolerance following disturbance in an eastern deciduous forest
• Alejandro A. Royo, Walter P. Carson. 2008. Direct and indirect effects of a dense understory on tree seedling recruitment in temperate forests: habitat-mediated predation versus competition
• Alejandro A. Royo. 2008. P.A. Thomas and J. Packham: Ecology of woodlands and forests: Description, dynamics, and diversity [book review]
• Alejandro A. Royo, Ramona M. Bates, Elizabeth P. Lacey. 2008. Demographic constraints in three populations of Lobelia boykinii, a rare wetland endemic
• Alejandro A. Royo, Walter P. Carson. 2006. On the formation of dense understory layers in forests worldwide: consequences and implications for forest dynamics, biodiversity, and succession
• Lauren Comisky, Alejandro A. Royo, Walter P. Carson. 2005. Deer Browsing Creates Rock Refugia Gardens on Large Boulders in the Allegheny National Forest, Pennsylvania
• Walter P. Carson, Joshua A. Banta, Alejandro A. Royo, Chad Kirschbaum. 2005. Plant communities growing on boulders in the Allegheny National Forest: Evidence for boulders as refugia from deer and as a bioassay of overbrowsing
• Kurt O. Reinhart, Alejandro A. Royo, Wim H. Van Der Putten, Keith Clay. 2005. Soil feedback and pathogen activity in Prunus serotina throughout its native range
• Alejandro A. Royo, Walter P. Carson. 2005. The herb community of a tropical forest in central Panama: dynamics and impact of mammalian herbivores

• Program
  

  Ecological Sustainability of Central Appalachian Forests

  https://research.fs.usda.gov/nrs/programs/appalachian
  Station

  Northern Research Station
• Experimental Forest or Range
  

  Kane Experimental Forest

  https://research.fs.usda.gov/nrs/forestsandranges/locations/kane
  Station

  Northern Research Station
• Project
  

  Desired REgeneration through Assisted Migration (DREAM)

  https://research.fs.usda.gov/nrs/projects/dream
  Station

  Northern Research Station
• Project
  

  Forest Management to Save the Moose in Minnesota

  https://research.fs.usda.gov/nrs/projects/minnesota-moose
  Station

  Northern Research Station

Multimedia

• Webinars

  Science x Wind Disturbance

  https://research.fs.usda.gov/products/multimedia/webinars/science-x-wind-disturbance
• Webinars

  Foundations of Assisted Migration

  https://research.fs.usda.gov/nrs/products/multimedia/webinars/foundations-assisted-migration

• University of Pittsburgh, Ph.D., Ecology and Evolutionary Biology, 2005
• University of North Carolina, Greensboro, M.S., Department of Biological Sciences, 1998
• University of North Carolina, Greensboro, B.A., Department of Biological Sciences, 1993
• University of Pittsburgh, Graduate Certificate, Latin American Studies, 2005