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Nathan P. P. Havill, PhD

Nathan P. P. Havill, PhD

Research Entomologist

Northern Research Station

0000-0002-4004-8266

nathan.p.havill@usda.gov

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Treesearch Publications

Kaka Tshering, Namgay Shacha, Yonten Norbu, Tshering, Sonam Tashi, Zephyr Zembrzuski, Elena Karlsen-Ayala, Nathan P. P. Havill. 2025. Diversity, distribution, biology, and predators of adelgids (Hemiptera: Adelgidae) in the conifer forests of Bhutan
Nikoleta Eleftheriadou, M. Lukas Seehausen, Marc Kenis, Greg Lefoe, Umar K Lubanga, Stephen D. Gaimari, Nathan P. P. Havill, Antonio P. Garonna, Nickolas G. Kavallieratos, Dimitrios N. Avtzis. 2025. Genetic diversity among Neoleucopis (Diptera: Chamaemyiidae) associated with Marchalina hellenica (Hemiptera: Marchalinidae) in Greece and Italy (Ischia)
Bryan M T Brunet, Dustin T Dial, Gaelen R Burke, Carol D von Dohlen, Julia Douglas Freitas, Haley Sanderson, Afiya R Chida, Samantha J Jones, Fergal J Martin, Leanne Haggerty, Stephen W Scherer, Ioannis Ragoussis, Steven J M Jones, Robert G Foottit, Nathan P. P. Havill. 2025. The Genome Sequence of the Hemlock Woolly Adelgid, Adelges tsugae Annand 1924
Katherine A. McManus, Therese M. M. Poland, Nathan P. P. Havill. 2025. Proceedings, 32nd USDA interagency research forum on invasive species
Tonya Bittner, Nathan P. P. Havill, Nicholas Dietschler, Zephyr Zembrzuski, Mark Whitmore, Norman Leppla. 2025. Eastern and western lineages of Leucotaraxis argenticollis (Diptera: Chamaemyiidae), a biological control agent for Adelges tsugae (Hemiptera: Adelgidae), are capable of hybridization in the laboratory
Ashley N. Schulz, Nathan P. P. Havill, Travis D. Marsico, Matthew P. Ayres, Kamal J.K. Gandhi, Daniel A. Herms, Angela M. Hoover, Ruth A. Hufbauer, Andrew M. Liebhold, Kenneth F. Raffa, Kathryn A. Thomas, Patrick Tobin, Daniel R. Uden, Angela M. Mech. 2025. What is a specialist? Quantifying host breadth enables impact prediction for invasive herbivores
Nathan P. P. Havill, Bryan Brunet, Zephyr Zembrzuski, Kaka Tshering. 2025. Four new adelgid (Hemiptera: Adelgidae) species from Bhutan, including the first legless species of Aphidomorpha
Katherine A. McManus, Therese M. M. Poland, Nathan P. P. Havill. 2025. Proceedings, 31st USDA interagency research forum on invasive species
Dustin Dial, Kathryn Weglarz, Bryan Brunet, Nathan P. P. Havill, Carol von Dohlen, Gaelen Burke. 2024. Whole-genome sequence of the Cooley spruce gall adelgid, Adelges cooleyi (Hemiptera: Sternorrhyncha: Adelgidae)
Soňa Šenfeldová, Thomas H. Atkinson, Miloš Knížek, Robert J. Rabaglia, Nathan P. P. Havill, Samuel F. Ward, Marek Turčáni, Andrew M. Liebhold. 2024. Determinants of host breadth in non-native bark and ambrosia beetles
Gwylim S. Blackburn, Christopher I Keeling, Julien Prunier, Melody Keena, Catherine Béliveau, Richard Hamelin, Nathan P. P. Havill, Francois Olivier Hebert, Roger C. Levesque, Michel Cusson, Ilga Porth. 2024. Genetics of flight in spongy moths (Lymantria dispar ssp.): functionally integrated profiling of a complex invasive trait
Bryan M.T. Brunet, Michael J. Raupach, Heinz-Otto Rehage, Nathan P. P. Havill, Robert G. Foottit. 2023. Discovery of the primary aphid (Hemiptera: Aphidomorpha) and scale insect (Hemiptera: Coccomorpha) type specimens from the collection of Theodor Hartig (1805–1880)
Nathan P. P. Havill, Shigehiko Shiyake, Zephyr Zembrzuski, Robert G. Foottit, Bryan M. T. Brunet. 2023. First record of Adelges ( Cholodkovskya ) viridanus (Hemiptera: Adelgidae) in North America, with a description of the fundatrix form in Japan
Jeremy Andersen, Nathan P. P. Havill, Jennifer L. Chandler, George Boettner, Brian Griffin, Joseph S. Elkinton. 2023. Seasonal differences in the timing of flight between the invasive winter moth and native Bruce spanworm promotes reproductive isolation
Albert (Bud) E. Mayfield, Tonya D. Bittner, Nicholas J. Dietschler, Joseph S. Elkinton, Nathan P. P. Havill, Melody Keena, David L. Mausel, James R. Rhea, Scott M. Salom, Mark C. Whitmore. 2023. Biological control of hemlock woolly adelgid in North America: History, status, and outlook.
Nathan P. P. Havill, Tonya Bittner, Jeremy Andersen, Nicholas Dietschler, Joseph Elkinton, Stephen Gaimari, Brian Griffin, Zephyr Zembrzuski, Mark C. Whitmore. 2023. Prey-associated genetic differentiation in two species of silver fly (Diptera: Chamaemyiidae), Leucotaraxis argenticollis and L. piniperda
Mathias Just Justesen, M. Lukas Seehausen, Nathan P. P. Havill, Marc Kenis, Stephen D. Gaimari, Izolda Matchutadze, Zephyr Zembrzuski, Hans Peter Ravn. 2023. Evaluation of Leucopis hennigrata (Diptera: Chamaemyiidae) as a classical biological control agent of Adelges nordmannianae (Hemiptera: Adelgidae) in northern Europe
Daniel R. Uden, Angela M. Mech, Nathan P. P. Havill, Ashley N. Schulz, Matthew P. Ayres, Daniel A. Herms, Angela M. Hoover, Kamal J. K. Gandhi, Ruth A. Hufbauer, Andrew M. Liebhold, Travis D. Marsico, Kenneth F. Raffa, Kathryn A. Thomas, Patrick Tobin, Craig R. Allen. 2022. Phylogenetic risk assessment is robust for forecasting the impact of European insects on North American conifers
Sabrina Celis, Nicholas Dietschler, Tonya Bittner, Nathan P. P. Havill, Michael Gates, Matthew Buffington, Mark C. Whitmore. 2022. Hymenopteran Parasitoids of Leucotaraxis argenticollis (Diptera: Chamaemyiidae) and Leucotaraxis piniperda : Implications for Biological Control of Hemlock Woolly Adelgid (Hemiptera: Adelgidae)
Nathan P. P. Havill, Stephen M. Dickson, Allison Kanoti, Michael S. Parisio, Jennifer Weimer, Zephyr Zembrzuski. 2022. Mass deposition of hemlock woolly adelgid sexuparae on New England beaches
Anish Kirtane, Nicholas J. Dietschler, Tonya D. Bittner, Marshall Bigler Lefebvre, Sabrina Celis, Katharine O'Connor, Nathan P. P. Havill, Mark C. Whitmore. 2022. Sensitive environmental DNA (eDNA) methods to detect hemlock woolly adelgid and its biological control predators Leucotaraxis silver flies and a Laricobius beetle
Jeremy C. Andersen, Nathan P. P. Havill, George H. Boettner, Jennifer L. Chandler, Adalgisa Caccone, Joseph S. Elkinton. 2022. Real‐time geographic settling of a hybrid zone between the invasive winter moth (Operophtera brumata L.) and the native Bruce spanworm (O. bruceata Hulst)
Dustin T. Dial, Kathryn M. Weglarz, Akintunde O. Aremu, Nathan P. P. Havill, Taylor A. Pearson, Gaelen R. Burke, Carol D. von Dohlen. 2022. Transitional genomes and nutritional role reversals identified for dual symbionts of adelgids (Aphidoidea: Adelgidae)
Stephen D. Gaimari, Nathan P. P. Havill. 2021. A new genus of Chamaemyiidae (Diptera: Lauxanioidea) predaceous on Adelgidae (Hemiptera), with a key to chamaemyiid species associated with Pinaceae-feeding Sternorrhyncha
Ashley N. Schulz, Angela M. Mech, Matthew P. Ayres, Kamal J. K. Gandhi, Nathan P. P. Havill, Daniel A. Herms, Angela M. Hoover, Ruth A. Hufbauer, Andrew M. Liebhold, Travis D. Marsico, Kenneth F. Raffa, Patrick Tobin, Daniel R. Uden, Kathryn A. Thomas. 2021. Predicting non-native insect impact: focusing on the trees to see the forest
Ryan C. Garrick, Ísis C. Arantes, Megan B. Stubbs, Nathan P. P. Havill. 2021. Weak spatial-genetic structure in a native invasive, the southern pine beetle (Dendroctonus frontalis), across the eastern United States
Jeremy C. Andersen, Nathan P. P. Havill, Brian P. Griffin, Jane U. Jepsen, Snorre B. Hagen, Tero Klemola, Isabel C. Barrio, Sofie A. Kjeldgaard, Toke T. Høye, John Murlis, Yuri N. Baranchikov, Andrey V Selikhovkin, Ole P.L. Vindstad, Adalgisa Caccone, Joseph S. Elkinton. 2021. Northern Fennoscandia via the British Isles: evidence for a novel post-glacial recolonization route by winter moth (Operophtera brumata)
Jeremy C. Andersen, Nathan P. P. Havill, Adalgisa Caccone, Joseph S. Elkinton. 2021. Four times out of Europe: Serial invasions of the winter moth, Operophtera brumata, to North America
Andrew M. Liebhold, Faith T. Campbell, Doria R. Gordon, Qinfeng Guo, Nathan P. P. Havill, Bradley Kinder, Richard A. MacKenzie, David R. Lance, Dean E. Pearson, Sharlene E. Sing, Travis Warziniack, Robert C. Venette, Denys Yemshanov. 2021. The Role of International Cooperation in Invasive Species Research
Nathan P. P. Havill, Brian P. Griffin, Jeremy C. Andersen, Robert G. Foottit, Mathias J. Justesen, Adalgisa Caccone, Vince D'Amico, Joseph S. Elkinton. 2021. Species delimitation and invasion history of the balsam woolly adelgid, Adelges ( Dreyfusia ) piceae (Hemiptera: Aphidoidea: Adelgidae), species complex
Brian Griffin, Jennifer L. Chandler, Jeremy Andersen, Nathan P. P. Havill, Joseph S. Elkinton. 2020. The Reliability of Genitalia Morphology to Monitor the Spread of the Invasive Winter Moth (Lepidoptera: Geometridae) in Eastern North America
Raymond J. Gagné, Nathan P. P. Havill. 2020. A New Species of Lestodiplosis (Diptera: Cecidomyiidae) Preying on Pineus (Hemiptera: Adelgidae), with a Redescription of Lestodiplosis juniperina (Felt)
Gina Davis, Laura Lowrey, Tom Eckberg, Amy Gannon, Danielle Malesky, Nathan P. P. Havill. 2020. Notes on balsam woolly adelgid, Adelges piceae (Ratzeburg, 1844) (Hemiptera: Adelgidae), range expansion in Idaho, Montana and Utah
Alex N. Neidermeier, Darrell W. Ross, Nathan P. P. Havill, Kimberly F. Wallin. 2020. Temporal Asynchrony of Adult Emergence Between Leucopis argenticollis and Leucopis piniperda (Diptera: Chamaemyiidae), Predators of the Hemlock Woolly Adelgid (Hemiptera: Adelgidae), with Implications for Biological Control
Alexander Rose, Darrell W. Ross, Nathan P. P. Havill, Kyle Motley, Kimberly F. Wallin. 2020. Coexistence of three specialist predators of the hemlock woolly adelgid in the Pacific Northwest USA
Laura Blackburn, Joseph S. Elkinton, Nathan P. P. Havill, Hannah J. Broadley, Jeremy C. Andersen, Andrew M. Liebhold. 2020. Predicting the invasion range for a highly polyphagous and widespread forest herbivore
Ashley N. Schulz, Angela M. Mech, Craig R. Allen, Matthew P. Ayres, Kamal J. K. Gandhi, Jessica Gurevitch, Nathan P. P. Havill, Daniel A. Herms, Ruth A. Hufbauer, Andrew M. Liebhold, Kenneth F. Raffa, Michael J. Raupp, Kathryn A. Thomas, Patrick C. Tobin, Travis D. Marsico. 2020. The impact is in the details: evaluating a standardized protocol and scale for determining non-native insect impact
Joseph S. Elkinton, Tonya D. Bittner, Valerie J. Pasquarella, George H. Boettner, Andrew M. Liebhold, Juli R. Gould, Heather Faubert, Lisa Tewksbury, Hannah J. Broadley, Nathan P. P. Havill, Ann E. Hajek. 2019. Relating Aerial Deposition of Entomophaga maimaiga Conidia (Zoopagomycota: Entomophthorales) to Mortality of Gypsy Moth (Lepidoptera: Erebidae) Larvae and Nearby Defoliation
Jeremy C. Andersen, Nathan P. P. Havill, Hannah J. Broadley, George H. Boettner, Adalgisa Caccone, Joseph S. Elkinton. 2019. Widespread hybridization among native and invasive species of Operophtera moths (Lepidoptera: Geometridae) in Europe and North America
Jeremy C. Andersen, Nathan P. P. Havill, Yaussra Mannai, Olfa Ezzine, Samir Dhahri, Mohamed Lahbib Ben Jamâa, Adalgisa Caccone, Joseph S. Elkinton. 2019. Identification of winter moth (Operophtera brumata) refugia in North Africa and the Italian Peninsula during the last glacial maximum
J.C. Andersen, C.P. Camp, M.J. Davis, Nathan P. P. Havill, J.S. Elkinton. 2019. Development of microsatellite markers for an outbreaking species of oak gall wasp, Zapatella davisae (Hymenoptera: Cynipidae), in the northeastern United States
Scott M. Salom, Gina Davis, Joseph Elkinton, Jeremiah Foley, Nathan P. P. Havill, Carrie Jubb, Albert Mayfield, Tom McAvoy, Rusty Rhea, R. Talbot Trotter, III, Mark Whitmore. 2019. A response to "Media representation of hemlock woolly adelgid management risks: a case study of science communication and invasive species control," published in biological invasions online on September 18, 2018
Nathan P. P. Havill, Anthony Cognato, Ek del-Val, Robert Rabaglia, Ryan C. Garrick. 2019. New Molecular Tools for Dendroctonus frontalis (Coleoptera: Curculionidae: Scolytinae) Reveal an East–West Genetic Subdivision of Early Pleistocene Origin
Tonya D. Bittner, Nathan P. P. Havill, Isis A.L. Caetano, Ann E. Hajek. 2019. Efficacy of Kamona strain Deladenus siricidicola nematodes for biological control of Sirex noctilio in North America and hybridisation with invasive conspecifics
Holly A. Wantuch, Nathan P. P. Havill, E. Richard Hoebeke, Thomas P. Kuhar, Scott M. Salom. 2019. Predators associated with the pine bark adelgid (Hemiptera: Adelgidae), a native insect in Appalachian forests, United States of America, in its southern range
Nicholas J. Russo, Chris S. Elphick, Nathan P. P. Havill, Morgan W. Tingley. 2019. Spring bird migration as a dispersal mechanism for the hemlock woolly adelgid
Angela M. Mech, Sherilyn J. Harper, Nathan P. P. Havill, Carol D. von Dohlen, Gaelen R. Burke. 2019. Ecological factors influencing the beneficial endosymbionts of the hemlock woolly adelgid (Hemiptera: Adelgidae)
Molly Darr, Rachel Brooks, Nathan P. P. Havill, E. Hoebeke, Scott Salom. 2018. Phenology and Synchrony of Scymnus coniferarum (Coleoptera: Coccinellidae) with Multiple Adelgid Species in the Puget Sound, WA, USA
Kathryn M. Weglarz, Nathan P. P. Havill, Gaelen R. Burke, Carol D. von Dohlen. 2018. Partnering With a Pest: Genomes of Hemlock Woolly Adelgid Symbionts Reveal Atypical Nutritional Provisioning Patterns in Dual-Obligate Bacteria
Molly Darr, Scott Salom, Rachel K. Brooks, Robert G. Foottit, Gary L. Miller, Nathan P. P. Havill. 2018. First report of Pineus strobi (Hartig, 1839) (Hemiptera: Adelgidae) in western North America
Sandrine Picq, Melody Keena, Nathan P. P. Havill, Don Stewart, Esther Pouliot, Brian Boyle, Roger C. Levesque, Richard C. Hamelin, Michel Cusson. 2018. Assessing the potential of genotyping-by-sequencing-derived single nucleotide polymorphisms to identify the geographic origins of intercepted gypsy moth (Lymantria dispar) specimens: A proof-of-concept study
Nathan P. P. Havill, Stephen D. Gaimari, Adalgisa Caccone. 2018. Cryptic east-west divergence and molecular diagnostics for two species of silver flies (Diptera: Chamaemyiidae: Leucopis ) from North America being evaluated for biological control of hemlock woolly adelgid
Garth Holman, Peter Del Tredici, Nathan P. P. Havill, Nam Sook Lee, Richard Cronn, Sarah Mathews, Linda Raubeson, Christopher S. Campbell. 2017. A new species and introgression in eastern Asian hemlocks (Pinaceae: Tsuga)
Jeremy C. Andersen, Nathan P. P. Havill, Adalgisa Caccone, Joseph S. Elkinton. 2017. Postglacial recolonization shaped the genetic diversity of the winter moth (Operophtera brumata) in Europe
Carol D. von Dohlen, Usha Spaulding, Kistie B. Patch, Kathryn M. Weglarz, Robert G. Foottit, Nathan P. P. Havill, Gaelen R. Burke. 2017. Dynamic Acquisition and Loss of Dual-Obligate Symbionts in the Plant-Sap-Feeding Adelgidae (Hemiptera: Sternorrhyncha: Aphidoidea)
Nathan P. P. Havill, J. Elkinton, J.C. Andersen, S.B. Hagen, Hannah J. Broadley, G.J. Boettner, A. Caccone. 2017. Asymmetric hybridization between non-native winter moth, Operophtera brumata (Lepidoptera: Geometridae), and native Bruce spanworm, Operophtera bruceata, in the Northeastern United States, assessed with novel microsatellites and SNPs
Nathan P. P. Havill, Shigehiko Shiyake, Ashley Lamb Galloway, Robert G. Foottit, Guoyue Yu, Annie Paradis, Joseph Elkinton, Michael Montgomery, Masakazu Sano, Adalgisa Caccone. 2016. Ancient and modern colonization of North America by hemlock woolly adelgid, Adelges tsugae (Hemiptera: Adelgidae), an invasive insect from East Asia
Colin Favret, Nathan P. P. Havill, Gary L. Miller, Masakazu Sano, Benjamin Victor. 2015. Catalog of the adelgids of the world (Hemiptera, Adelgidae)
Melissa J. Fischer, Carlyle C. Brewster, Nathan P. P. Havill, Scott M. Salom, Loke T. Kok. 2015. Assessment of the potential for hybridisation between Laricobius nigrinus (Coleoptera: Derodontidae) and Laricobius osakensis, predators of the hemlock woolly adelgid (Hemiptera: Adelgidae)
Melissa J. Fischer, Nathan P. P. Havill, Carlyle C. Brewster, Gina A. Davis, Scott M. Salom, Loke T. Kok. 2015. Field assessment of hybridization between Laricobius nigrinus and L. rubidus, predators of Adelgidae
Michael Montgomery, Nathan P. P. Havill. 2014. Balsam woolly adelgid
C.E. Jones, Nathan P. P. Havill, James Hanula, S.K. Braman. 2014. Post release recovery of hemlock woolly adelgid predators in the North Georgia mountains.
Albert (Bud) E. Mayfield, Barbara C. Reynolds, Carla I. Coots, Nathan P. P. Havill, Cavell Brownie, Andrew R. Tait, James Hanula, Shimat V. Joseph, Ashley B. Galloway. 2014. Establishment, hybridization and impact of Laricobius predators on insecticide-treated hemlocks: Exploring integrated management of the hemlock woolly adelgid
Nathan P. P. Havill, Ligia C. Vieira, Scott M. Salom. 2014. Hemlock woolly adelgid: a threat to eastern forests
Nathan P. P. Havill, Ligia C. Vieira, Scott M. Salom. 2014. Biology and control of hemlock woolly adelgid
Melissa J. Fischer, Nathan P. P. Havill, Carrie S. Jubb, Sean W. Prosser, Brent D. Opell, Scott M. Salom, Loke T. Kok. 2014. Contamination delays the release of Laricobius osakensis for biological control of hemlock woolly adelgid: cryptic diversity in Japanese Laricobius spp
Stephen P. Cook, Frank Merickel, Karen Ward, Nathan P. P. Havill. 2013. First report of Adelges abietis (Linneaus) (Hemiptera: Adelgidae) in Idaho
H.P. Ravn, Nathan P. P. Havill, S. Akbulut, R.G. Foottit, M. Serin, M. Erdem, S. Mutun, M. Kenis. 2013. Dreyfusia nordmannianae in Northern and Central Europe: potential for biological control and comments on its taxonomy
Nathan P. P. Havill, Gina Davis, David Mausel, Joanne Klein, Richard McDonald, Cera Jones, Melissa Fischer, Scott Salom, Adelgisa Caccone. 2012. Hybridization between a native and introduced predator of Adelgidae: An unintended result of classical biological control
G.A. Davis, Nathan P. P. Havill, Z.N. Adelman, A. Caccone, L.T. Kok, S.M. Salom. 2011. DNA barcodes and molecular diagnostics to distinguish an introduced and native Laricobius (Coleoptera: Derodontidae) species in eastern North America
Nathan P. P. Havill, Scott Salom, Gina Davis, Melissa Fischer, David Mausel, Bradley Onken. 2011. The introduction of Laricobius nigrinus as a biological control agent for the hemlock woolly adelgid: Is there a threat to the native congener, L. rubidus?
Nathan P. P. Havill, Michael Montgomery, Melody Keena. 2011. Hemlock woolly adelgid and its hemlock hosts: A global perspective
Michael Montgomery, S. Shiyake, Nathan P. P. Havill. 2011. A new species of Laricobius (Coleoptera: Derodontidae) from Japan with phylogeny and a key for native and introduced congeners in North America
Nathan P. P. Havill, Gina Davis, Joanne Klein, Adalgisa Caccone, Scott Salom. 2011. Hemlock woolly adelgid biological control: molecular methods to distinguish Laricobius nigrinus, L. rubidus, and their hybrids
Masakazu Sano, Nathan P. P. Havill, Kenichi Ozaki. 2011. Taxonomic identity of a galling adelgid (Hemiptera: Adelgidae) from three spruce species in central Japan
Zi-xiang Yang, Xiao-ming Chen, Nathan P. P. Havill, Ying Feng, Hang Chen. 2010. Phylogeny of Rhus gall aphids (Hemiptera:Pemphigidae) based on combined molecular analysis of nuclear EF1α and mitochondrial COII genes
Nathan P. P. Havill, Michael Montgomery, Shigehiko Shiyake, Ashley Lamb, Melody Keena, Adalgisa Caccone. 2009. Hemlock woolly adelgid population genetics
Nathan P. P. Havill, Joanne Klein, Melody Keena, Adalgisa Caccone. 2009. Characterization of microsatellite loci for Laricobius nigrinus and L rubidus, predators of adelgids in North America
R.G. Foottit, H.E.L. Maw, Nathan P. P. Havill, R.G. Ahern, Michael Montgomery. 2009. DNA barcodes to identify species and explore diversity in the Adelgidae (Insecta: Hemiptera: Aphidoidea)
Nathan P. P. Havill, Michael Montgomery. 2008. The role of arboreta in studying the evolution of host resistance to the hemlock woolly adelgid
Michael Montgomery, Nathan P. P. Havill, Carole Cheah, Mark McClure, Gabriella Zilahi-Balogh, Ashley Lamb, Scott Salom. 2003. Hemlock woolly adelgid biological control research
Elizabeth Butin, Joseph Elkinton, Nathan P. P. Havill, Michael Montgomery. 2003. Comparative impact of Scymnus ningshanensis and Pseudoscymnus tsugae (Coleoptera: Coccinellidae) on the hemlock woolly adelgid
Elizabeth Butin, Michael Montgomery, Nathan P. P. Havill, Joseph Elkinton. 2002. Pre-release host range assessment for classical biological controls: experience with predators for the hemlock woolly adelgid
Michael Montgomery, Hongbin Wang, Defu Yao, Wenhau Lu, Nathan P. P. Havill, Guangwu Li. 2002. Biology of Scymnus ningshanensis (Coleoptera: Coccinellidae): A predator of Adelges tsugae (Homoptera: Adelgidae)

Yale University, Ph.D., Ecology & Evolutionary Biology, 2006
Yale University, M.S., Ecology & Evolutionary Biology, 2003
University of Wisconsin, Madison, M.S., Entomology, 1998
College of William and Mary, B.S., Biology, 1996

Last updated March 10, 2026

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