Sarah Jovan, PhD

Research Ecologist

Pacific Northwest Research Station

0000-0001-7860-4005

sarah.jovan@usda.gov

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Treesearch Publications

Sian Kou-Giesbrecht, Carla R. Reis Ely, Steven S. Perakis, Cory C. Cleveland, Duncan N. L. Menge, Sasha C. Reed, Benton N. Taylor, Sarah A. Batterman, Timothy E. Crews, Katherine A. Dynarski, Maga Gei, Michael J. Gundale, David F. Herridge, Sarah Jovan, Mark B. Peoples, Johannes Piipponen, Emilio Rodríguez-Caballero, Verity G. Salmon, Fiona M. Soper, Anika P. Staccone, Bettina Weber, Christopher A. Williams, Nina Wurzburger. 2025. Overestimated natural biological nitrogen fixation translates to an exaggerated CO2 fertilization effect in Earth system models
Carla R. Reis Ely, Steven S. Perakis, Cory C. Cleveland, Duncan N. L. Menge, Sasha C. Reed, Sarah A. Batterman, Timothy E. Crews, Katherine A. Dynarski, Maga Gei, Michael J. Gundale, Sarah Jovan, Sian Kou-Giesbrecht, Mark B. Peoples, Emilio Rodríguez-Caballero, Verity G. Salmon, Fiona M. Soper, Anika P. Staccone, Benton N. Taylor, Bettina Weber, Nina Wurzburger. 2025. A global dataset of terrestrial biological nitrogen fixation
Adrienne Kovasi, Bruce McCune, Sarah Jovan. 2025. Extracting gradients from community data with multiple stressors and empty plots: A case study and simulations
Christopher Zuidema, Michael Paulsen, Christopher D. Simpson, Sarah Jovan. 2024. Evaluation of Orthotrichum lyellii moss as a biomonitor of diesel exhaust
Sarah Jovan, Eleonore Jacobson, Jason M. Unrine, Nasser Jalili-Jahani, Bruce McCune. 2024. Putting biomonitors to work: native moss as a screening tool for solid waste incineration
Alyssa E. Shiel, Sarah Jovan, Christina J. Murphy. 2024. Lead-sheathed telecom cables and historic leaded gasoline emissions substantially raise environmental lead levels in Portland, Oregon
Monika Derrien, Weston Brinkley, Dale Blahna, Alberto Rodríguez, Roseann Barnhill, Christopher Zuidema, Katie Beaver, Elisabeth Grinspoon, Sarah Jovan. 2024. Joining collective impact and community science: a framework for core collaborative community science
Adrienne Kovasi, Bruce McCune, Sarah Jovan. 2023. Evaluating Letharia vulpina transplants for bioindication of nitrogen deposition
Susan Will-Wolf, Sarah Jovan. 2023. The National Lichen Atlas: summarizing 23 years of epiphytic lichen community data
Tara Greaver, Stephen McDow, Jennifer Phelan, S. Douglas Kaylor, Jeffrey D. Herrick, Sarah Jovan. 2023. Synthesis of lichen response to gaseous nitrogen: Ammonia versus nitrogen dioxide
John Kirkland, Sarah Jovan, Monika Derrien. 2023. The more the mossier: Using community science to map air quality in environmental justice investigations
Sarah Jovan, Christopher Zuidema, Monika Derrien, Amanda L. Bidwell, Weston Brinkley, Robert J. Smith, Dale Blahna, Roseann Barnhill, Linn Gould, Alberto J. Rodríguez, Michael C. Amacher, Troy D. Abel, Paulina López. 2022. Heavy metals in moss guide environmental justice investigation: A case study using community science in Seattle, WA, USA
Adrienne Kovasi, Bruce McCune, Sarah Jovan. 2022. The potential use of elemental content of saxicolous lichens as bioindicators of nitrogen deposition in the central and southern California mountains
Igor Burstyn, Geoffrey Donovan, Yvonne L. Michael, Sarah Jovan. 2022. Association of polycyclic aromatic hydrocarbons in moss with blood biomarker among nearby residents in Portland, Oregon
Bruce McCune, Sunia Yang, Sarah Jovan, Heather T. Root. 2022. Climate and epiphytic macrolichen communities in the Four Corners region of the U.S.A.
Michelle Kondo, Christopher Zuidema, Hector A. Moran, Sarah Jovan, Monika Derrien, Weston Brinkley, Anneclaire J. De Roos, Loni Philip. Tabb. 2022. Spatial predictors of heavy metal concentrations in epiphytic moss samples in Seattle, WA
Sarah Jovan, Michael Haldeman, Susan Will-Wolf, Karen Dillman, Linda Geiser, Joel Thompson, Daphne Stone, Jason Hollinger. 2021. National atlas of epiphytic lichens in forested habitats of the United States
Sarah Jovan, Mark E. Fenn, Monica Buhler, Andrzej Bytnerowicz, Adrienne Kovasi, Martin Hutten, Elisa DiMeglio, Donald Schweizer. 2021. Challenges characterizing N deposition to high elevation protected areas: A case study integrating instrument, simulated, and lichen inventory datasets for the Devils Postpile National Monument and surrounding region, USA
Anthony W. D'Amato, Sarah Jovan, Christopher Looney. 2021. Investigating linkages between the size-growth relationship and drought, nitrogen deposition, and structural complexity in western U.S. Forests
Heather T. Root, Sarah Jovan, Mark E. Fenn, Michael Amacher, Josh Hall, John D. Shaw. 2021. Lichen bioindicators of nitrogen and sulfur deposition in dry forests of Utah and New Mexico, USA
Sarah Jovan, Susan Will-Wolf, Linda Geiser, Karen Dillman, Michael Haldeman, Rick Shory. 2020. User guide for the national Forest Inventory and Analysis lichen database (Version 1.0)
Monika Derrien, Christopher Zuidema, Sarah Jovan, Amanda Bidwell, Weston Brinkley, Paulina López, Roseann Barnhill, Dale Blahna. 2020. Toward environmental justice in civic science: Youth performance and experience measuring air pollution using moss as a bio-indicator in industrial-adjacent neighborhoods
Robert J. Smith, Sarah Jovan, Bruce McCune. 2020. Climatic niche limits and community‐level vulnerability of obligate symbioses
Mark E. Fenn, Haiganoush K. Preisler, Jeremy S. Fried, Andrzej Bytnerowicz, Susan L. Schilling, Sarah Jovan, Olaf Kuegler. 2020. Evaluating the effects of nitrogen and sulfur deposition and ozone on tree growth and mortality in California using a spatially comprehensive forest inventory
Susan Will-Wolf, Sarah Jovan, Michael C. Amacher, Paul L. Patterson. 2020. Lichen elemental indicators for air pollution in Eastern United States forests; a pilot study in the upper Midwest
Linda H. Geiser, Peter R. Nelson, Sarah Jovan, Heather T. Root, Christopher M. Clark. 2019. Assessing ecological risks from atmospheric deposition of nitrogen and sulfur to US forests using epiphytic macrolichens
Christopher M. Clark, Samuel M. Simkin, Edith B. Allen, William D. Bowman, Jayne Belnap, Matthew L. Brooks, Scott L. Collins, Linda H. Geiser, Frank S. Gilliam, Sarah Jovan, Linda Pardo, Bethany K. Schulz, Carly J. Stevens, Katharine N. Suding, Heather L. Throop, Donald M. Waller. 2019. Potential vulnerability of 348 herbaceous species to atmospheric deposition of nitrogen and sulfur in the United States
Robert Smith, Peter R. Nelson, Sarah Jovan, Paul J. Hanson, Bruce McCune. 2018. Novel climates reverse carbon uptake of atmospherically dependent epiphytes: Climatic constraints on the iconic boreal forest lichen Evernia mesomorpha
Robert Pattison, Hans Andersen, Andrew Gray, Bethany K. Schulz, Robert Smith, Sarah Jovan. 2018. Forests of the Tanana Valley State Forest and Tetlin National Wildlife Refuge, Alaska: results of the 2014 pilot inventory
Susan Will-Wolf, Sarah Jovan, Michael C. Amacher. 2018. Chapter 6 - Lichen species to bioindicate air quality in Eastern United States from elemental composition: lessons from the Midwest
Natasha Vizcarra, Sarah Jovan, Demetrios Gatziolis, Vicente Monleon. 2018. Of moss and men: Using moss as a bioindicator of toxic heavy metals at the city scale
Sheel Bansal, Leslie C. Brodie, Sharon Stanton, Karen Waddell, Marin Palmer, Glenn Christensen, Olaf Kuegler, John M. Chase, Joel Thompson, Sarah Jovan, Andrew Gray, Morgan Todd. 2017. Oregon’s forest resources, 2001–2010: ten-year Forest Inventory and Analysis report
Robert J. Smith, Sarah Jovan, Andrew Gray, Bruce McCune. 2017. Sensitivity of carbon stores in boreal forest moss mats - effects of vegetation, topography and climate
Susan Will-Wolf, Sarah Jovan, Michael C. Amacher. 2017. Lichen elemental content bioindicators for air quality in upper Midwest, USA: A model for large-scale monitoring
Susan Will-Wolf, Sarah Jovan, Michael C. Amacher. 2017. Lichen elements as pollution indicators: evaluation of methods for large monitoring programmes
Robert J. Smith, Sarah Jovan, Bruce McCune. 2017. Lichen communities as climate indicators in the U.S. Pacific States.
Therese S. Carter, Christopher M. Clark, Mark E. Fenn, Sarah Jovan, Steven S. Perakis, Jennifer Riddell, Paul Schaberg, Tara L. Greaver, Meredith G. Hastings. 2017. Mechanisms of nitrogen deposition effects on temperate forest lichens and trees
Samuel M. Simkin, Edith B. Allen, William D. Bowman, Christopher M. Clark, Jayne Belnap, Matthew L. Brooks, Brian S. Cade, Scott L. Collins, Linda H. Geiser, Frank S. Gilliam, Sarah Jovan, Linda Pardo, Bethany K. Schulz, Carly J. Stevens, Katharine N. Suding, Heather L. Throop, Donald M. Waller. 2016. Conditional vulnerability of plant diversity to atmospheric nitrogen deposition across the United States
Demetrios Gatziolis, Sarah Jovan, Geoffrey Donovan, Michael Amacher, Vicente Monleon. 2016. Elemental atmospheric pollution assessment via moss-based measurements in Portland, Oregon
Charles E. Werstak, John D. Shaw, Sara A. Goeking, Christopher Witt, James S. Menlove, Mike T. T. Thompson, R. Justin DeRose, Michael C. Amacher, Sarah Jovan, Todd A. Morgan, Colin B. Sorenson, Steven W. Hayes, Chelsea P. McIver. 2016. Utah's forest resources, 2003-2012
Geoffrey Donovan, Sarah Jovan, Demetrios Gatziolis, Igor Burstyn, Yvonne L. Michael, Michael C. Amacher, Vicente Monleon. 2016. Using an epiphytic moss to identify previously unknown sources of atmospheric cadmium pollution
Dominick A. DellaSala, Patric Brandt, Marni Koopman, Jessica Leonard, Claude Meisch, Patrick Herzog, Paul Alaback, Michael I. Goldstein, Sarah Jovan, Andy MacKinnon, Henrik von Wehrden. 2015. Climate change may trigger broad shifts in North America's Pacific Coastal rainforests
Geoffrey Donovan, Sarah Jovan, Demetrios Gatziolis, Vicente Monleon. 2015. Moss as bio-indicators of human exposure to polycyclic aromatic hydrocarbons in Portland, OR
Sarah Jovan, Susan Will-Wolf, Michael Amacher. 2015. “Lichens Lite?” chemical analysis of lichens for tracking 26 pollutants
Mark E. Fenn, Jeremy S. Fried, Haiganoush K. Preisler, Andrzej Bytnerowicz, Susan L. Schilling, Sarah Jovan, Olaf Kuegler. 2015. Remeasured FIA plots reveal tree-level diameter growth and tree mortality impacts of nitrogen deposition on California’s forests
Robert Smith, Sarah Jovan, Bruce McCune. 2015. Evaluating carbon stores at the earth-atmosphere interface: moss and lichen mats of subarctic Alaska
Linda S. Heath, Sarah M. Anderson, Marla R. Emery, Jeffrey A. Hicke, Jeremy Littell, Alan Lucier, Jeffrey G. Masek, David L. Peterson, Richard Pouyat, Kevin M. Potter, Guy Robertson, Jinelle Sperry, Andrzej Bytnerowicz, Sarah Jovan, Miranda H. Mockrin, Robert C. Musselman, Bethany K. Schulz, Robert Smith, Susan I. Stewart. 2015. Indicators of climate impacts for forests: recommendations for the US National Climate Assessment indicators system
David J. Nowak, Sarah Jovan, Christina Branquinho, Sofia Augusto, Manuel C. Ribeiro, Conor E. Kretsch. 2015. Biodiversity, air quality and human health
Sarah Jovan, Robert Smith, Juan C. Benavides, Michael Amacher, Bruce McCune. 2015. A rapid method for landscape assessment of carbon storage and ecosystem function in moss and lichen ground layers
Susan Will-Wolf, Sarah Jovan, Peter Neitlich, JeriLynn E. Peck, Roger Rosentreter. 2015. Lichen-based indices to quantify responses to climate and air pollution across northeastern U.S.A
Heather T. Root, Linda H. Geiser, Sarah Jovan, Peter Neitlich. 2015. Epiphytic macrolichen indication of air quality and climate in interior forested mountains of the Pacific Northwest, USA
Sarah Jovan. 2014. Chapter 7 - Climate effects on lichen indicators for nitrogen
Susan Will-Wolf, Randall Morin, Mark J. Ambrose, Kurt Riitters, Sarah Jovan. 2014. Chapter 6 - Links between land cover and lichen species richness at large scales in forested ecosystems across the United States.
Heather T. Root, Bruce McCune, Sarah Jovan. 2014. Lichen communities and species indicate climate thresholds in southeast and south-central Alaska, USA
Mark E. Fenn, H.-D. Nagel, I. Koseva, J. Aherne, Sarah Jovan, L.H. Geiser, A. Schlutow, T. Scheuschner, Andrzej Bytnerowicz, B.S. Gimeno, F. Yuan, S.A. Watmough, E.B. Allen, R.F. Johnson, T. Meixner. 2014. A comparison of empirical and modelled nitrogen critical loads for Mediterranean forests and shrublands in California
Heather T. Root, Linda H. Geiser, Mark E. Fenn, Sarah Jovan, Martin A. Hutten, Suraj Ahuja, Karen Dillman, David Schirokauer, Shanti Berryman, Jill A. McMurray. 2013. A simple tool for estimating throughfall nitrogen deposition in forests of western North America using lichens
Jill A. McMurray, Dave W. Roberts, Mark E. Fenn, Linda H. Geiser, Sarah Jovan. 2013. Using epiphytic lichens to monitor nitrogen deposition near natural gas drilling operations in the Wind River Range, WY, USA
Sarah Jovan. 2012. Lichens
Sarah Jovan. 2012. Epiphytic Macrolichen Community Composition Database—epiphytic lichen synusiae in forested areas of the US
Sarah Jovan, Jennifer Riddell, Pamela E. Padgett, Thomas Nash. 2012. Eutrophic lichens respond to multiple forms of N: implications for critical levels and critical loads research
Linda Geiser, Sarah Jovan, Marie Oliver. 2011. Canaries in a coal mine: using lichens to measure nitrogen pollution
Jennifer Riddell, Sarah Jovan, Pamela E. Padgett, Ken Sweat. 2011. Tracking lichen community composition changes due to declining air quality over the last century: the Nash legacy in Southern California
Christopher W. W. Woodall, Michael C. Amacher, William Bechtold, John W. Coulston, Sarah Jovan, Charles H. (Hobie) Perry, KaDonna Randolph, Bethany K. Schulz, Gretchen C. Smith, Susan Will-Wolf. 2011. Status and future of the forest health indicators program of the USA
Linda H. Geiser, Sarah Jovan, Doug A. Glavich, Matthew K. Porter. 2010. Lichen-based critical loads for atmospheric nitrogen deposition in Western Oregon and Washington forests, USA
Christopher W. W. Woodall, Barbara Conkling, Michael C. Amacher, John W. Coulston, Sarah Jovan, Charles H. (Hobie) Perry, Bethany K. Schulz, Gretchen C. Smith, Susan Will Wolf. 2010. The Forest Inventory and Analysis Database Version 4.0: Database Description and Users Manual for Phase 3
Sarah Jovan. 2009. Baseline results from the Lichen Community Indicator Program in the Pacific Northwest: Air quality patterns and evidence of a nitrogen pollution problem
Susan Will-Wolf, Sarah Jovan. 2009. Lichens, ozone, and forest health - exploring cross-indicator analyses with FIA data
Bruce McCune, Sarah Jovan, Amanda Hardman. 2008. Changes in forage lichen biomass after insect outbreaks and fuel reduction treatment in the Blue Mountains, Oregon
Mark E. Fenn, Sarah Jovan, F. Yuan, L. Geiser, T. Meixner, B.S. Gimeno. 2008. Empirical and simulated critical loads for nitrogen deposition in California mixed conifer forests
Sarah Jovan, Tom Carlberg. 2007. Nitrogen content of Letharia vulpina tissue from forests of the Sierra Nevada, California: geographic patterns and relationships to ammonia estimates and climate
Sarah Jovan, Bruce Mccune. 2006. Using epiphytic macrolichen communities for biomonitoring ammonia in forests of the greater Sierra Nevada, California
Sarah Jovan, Bruce McCune. 2005. Air-quality bioindication in the greater central valley of California, with epiphytic macrolichen communities.
Sarah Jovan, Bruce McCune. 2004. Regional variation in epiphytic macrolichen communities in northern and central California forests

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Oregon State University, PhD, Botany and Ecology, 2005

Last updated July 6, 2026

Sarah Jovan, PhD

Treesearch Publications

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