Anna W. Schoettle, PhD
Emeritus Research Plant Ecophysiologist
anna.schoettle@usda.gov
Treesearch Publications
- Sparkle L. Malone, Anna W. Schoettle, Kelly S. Burns, Holly S. J. Kearns, Jane E. Stewart, Maria Newcomb, Christy M. Cleaver. 2025. RustMapper: White pine blister rust risk across high elevation forests in the western United States
- Jun-Jun Liu, Richard A. Sniezko, Sydney Houston, Genoa Alger, Jodie Krakowski, Anna W. Schoettle, Robert Sissons, Arezoo Zamany, Holly Williams, Benjamin Rancourt, Angelia Kegley. 2025. A new threat to limber Pine (Pinus flexilis) restoration in Alberta and beyond: First documentation of a Cronartium ribicola race (vcr4) virulent to Cr4-controlled major gene resistance
- Paula J. Fornwalt, Andreas P. Wion, Anna W. Schoettle, Miranda D. Redmond, Steven Alton, Javier E. Mercado, Brice Hanberry, Jose Negron. 2024. Vulnerability of major Colorado Front Range tree species to climate change (Chapter 5)
- Lauren Hines-Acosta, Kelly Burns, Anna W. Schoettle, Jane Stewart, Wade Tinkham. 2024. Promoting healthy forests: Limber pine in the Rockies is vulnerable to long-term decline
- Kelly Burns, Erin Hooten, Holly Kearns, Maria Newcomb, Anna W. Schoettle, Erin Shanahan. 2023. Terminal branch dieback along Commissary Ridge in the Bridger-Teton National Forest: Investigating white pine blister rust canker ages and rates of expansion
- Olga Kozhar, Kelly S. Burns, Anna W. Schoettle, Jane E. Stewart. 2023. Distribution of cronartium x flexili, an interspecific hybrid of two fungal tree rust pathogens, in subalpine forest ecosystems of western USA
- Anna W. Schoettle. 2023. Mountain top sentinels
- Josh McDaniel, Anna W. Schoettle, Barbara J. Bentz, Sara A. Goeking, Robert E. Keane, Melissa Jenkins, Diana Tomback. 2023. Not on our watch: Scientists and managers rush to save high-elevation, five-needle pines
- Kelly S. Burns, Wade Tinkham, K. A. Leddy, Anna W. Schoettle, William R. Jacobi, Jane E. Stewart. 2023. Interactions between white pine blister rust, bark beetles, and climate over time indicate vulnerabilities to limber pine health
- Diana F. Tomback, Robert E. Keane, Anna W. Schoettle, Richard A. Sniezko, Melissa B. Jenkins, Cara R. Nelson, Andrew D. Bower, Clay R. DeMastus, Emily Guiberson, Jodie Krakowski, Michael P. Murray, Elizabeth R. Pansing, Julee Shamhart. 2022. Tamm review: Current and recommended management practices for the restoration of whitebark pine (Pinus albicaulis Engelm.), an imperiled high-elevation Western North American forest tree
- Jessica W. Wright, Melissa B. Jenkins, Anna W. Schoettle, Karl A. Anderson, Joseph Fortier, Linh Hoang, Tony Incashola, Robert E. Keane, Jodie Krakowski, Dawn M. LaFleur, Sabine Mellmann-Brown, Elliott D. Meyer, ShiNaasha Pete, Katherine Renwick, Robert A. Sissons. 2022. Restoring a forest keystone species: A plan for the restoration of whitebark pine (Pinus albicaulis Engelm.) in the Crown of the Continent ecosystem
- Jun-Jun Liu, Anna W. Schoettle, Richard A. Sniezko, Kristen M. Waring, Holly Williams, Arezoo Zamany, Jeremy S. Johnson, Angelia Kegley. 2022. Comparative association mapping reveals conservation of major gene resistance to white pine blister rust in southwestern white pine (Pinus strobiformis) and limber pine (P. flexilis)
- Anna W. Schoettle, Kelly S. Burns, Shawn McKinney, Jodie Krakowski, Kristen M. Waring, Diana F. Tomback, Marianne Davenport. 2022. Integrating forest health conditions and species adaptive capacities to infer future trajectories of the high elevation five-needle white pines
- Anna W. Schoettle, Angelia Kegley, Richard A. Sniezko, Kelly S. Burns, Detlev R. Vogler, Phyllis Pineda Bovin, Gretchen Baker. 2022. Preparing for invasion: Rust resistance in limber, Great Basin bristlecone, and Rocky Mountain bristlecone pines
- Robert E. Keane, Anna W. Schoettle, Diana F. Tomback. 2022. Effective actions for managing resilient high elevation five-needle white pine forests in western North America at multiple scales under changing climates
- Jun-Jun Liu, Anna W. Schoettle, Richard A. Sniezko, Holly Williams, Arezoo Zamany, Benjamin Rancourt. 2021. Fine dissection of limber pine resistance to Cronartium ribicola using targeted sequencing of the NLR family
- Mitra Menon, Justin C. Bagley, Gerald F. M. Page, Amy V Whipple, Anna W. Schoettle, Christopher J. Still, Christian Wehenkel, Kristen M. Waring, Lluvia Flores-Renteria, Samuel A. Cushman, Andrew J. Eckert. 2021. Adaptive evolution in a conifer hybrid zone is driven by a mosaic of recently introgressed and background genetic variants
- Kevin S. McKelvey, William M. Block, Terrie B. Jain, Charlie H. Luce, Deborah S. Page-Dumroese, Bryce A. Richardson, Vicki A. Saab, Anna W. Schoettle, Carolyn H. Sieg, Daniel R. Williams. 2021. Adapting research, management, and governance to confront socioecological uncertainties in novel ecosystems
- Jun-Jun Liu, Richard A. Sniezko, Robert Sissons, Jodie Krakowski, Genoa Alger, Anna W. Schoettle, Holly Williams, Arezoo Zamany, Rachel A. Zitomer, Angelia Kegle. 2020. Association mapping and development of marker-assisted selection tools for the resistance to white pine blister rust in the Alberta limber pine populations
- Anna W. Schoettle. 2020. Genetic resistance to white pine blister rust is key but how it is managed will determine the future of high elevation five-needle pines
- Melissa Jenkins, Tony Incashola, Dawn LaFleur, Linh Hoang, Katie Renwick, Anna W. Schoettle, Sabine Mellman-Brown, James Lozeau, Kelly Rose O’Bennick, Elliott Meyer, Karl Anderson, Rob Sissons, Robert E. Keane, Brad Jones. 2020. Crown of the Continent Ecosystem Whitebark Pine Restoration Strategy: 2019 Pilot - Project Summary
- Jun-Jun Liu, Anna W. Schoettle, Richard A. Sniezko. 2020. Development and application of high density genetic map of limber pine (Pinus Flexilis James) for genomics-based breeding
- Anna W. Schoettle, Kelly S. Burns, Christy M. Cleaver, Brian Verhulst, J. Jeff. Connor. 2020. Proactive limber pine conservation strategy for the Greater Rocky Mountain National Park Area - an overview
- James R. Meldrum, Patricia A. Champ, Craig Bond, Anna W. Schoettle. 2020. Paired stated preference methods for valuing management of white pine blister rust: Order effects and outcome uncertainty
- Jun-Jun Liu, Yu Xiang, Richard A. Sniezko, Anna W. Schoettle, Holly Williams, Arezoo Zamany. 2019. Characterization of Cronartium ribicola dsRNAs reveals novel members of the family Totiviridae and viral association with fungal virulence
- Anna W. Schoettle. 2019. Taking the long view and acting now - prioritizing management of high elevation five-needle pines
- Mitra Menon, Erin Landguth, Alejandro Leal‐Saenz, Justin C. Bagley, Anna W. Schoettle, Christian Wehenkel, Lluvia Flores‐Renteria, Samuel A. Cushman, Kristen M. Waring, Andrew J. Eckert. 2019. Tracing the footprints of a moving hybrid zone under a demographic history of speciation with gene flow
- Jun-Jun Liu, Anna W. Schoettle, Richard A. Sniezko, Fupan Yao, Arezoo Zamany, Holly Williams, Benjamin Rancourt. 2019. Limber pine (Pinus flexilis James) genetic map constructed by exome-seq provides insight into the evolution of disease resistance and a genomic resource for genomics-based breeding
- Anna W. Schoettle, Kelly S. Burns, Christy M. Cleaver, J. Jeff Connor. 2019. Proactive limber pine conservation strategy for the Greater Rocky Mountain National Park Area
- Anna W. Schoettle, William R. Jacobi, Kristen M. Waring, Kelly S. Burns. 2019. Regeneration for resilience framework to support regeneration decisions for species with populations at risk of extirpation by white pine blister rust
- Sparkle L. Malone, Anna W. Schoettle, Jonathan D. Coop. 2018. The future of subalpine forests in the Southern Rocky Mountains: Trajectories for Pinus aristata genetic lineages
- Anna W. Schoettle. 2018. What is in your planting bag?
- Christine T. Holtz, Anna W. Schoettle. 2018. Is resistance to mountain pine beetle associated with genetic resistance to white pine blister rust in limber pine?
- Anna W. Schoettle, K. S. Burns, R. A. Sniezko. 2018. The proactive strategy: Preparing the landscape for invasion by accelerating the evolution of resistance
- Anna W. Schoettle, R. A. Sniezko, A. Kegley, K. S. Burns. 2018. Patterns of resistance to Cronartium ribicola in Rocky Mountain bristlecone pine (Pinus aristata)
- Anna W. Schoettle, R. A. Sniezko, A. Kegley, K. S. Burns. 2018. Genetic resistance to Cronartium ribicola in limber pine (Pinus flexilis)
- Anna W. Schoettle, Betsy Goodrich, Jen Klutsch, Kelly Burns. 2018. White pine blister rust confirmed on limber pine (Pinus flexilis) in Rocky Mountain National Park
- Anna W. Schoettle, Kelly S. Burns, William Jacobi, John B. Popp, Sara Alberts, Tim Douville, Frank Romaro. 2018. Southern Rockies rust resistance trial
- Anna W. Schoettle, J. G. Klutsch, M. F. Antolin, S. Field. 2018. Population projections and resistance gene frequencies in the presence of white pine blister rust: A population genetic model for high elevation five-needle pines
- Patrick I. Bennett, Jerry Hill, Douglas P. Savin, Angelia Kegley, Anna W. Schoettle, Richard A. Sniezko, Ben Bird, Jeffrey Stone. 2018. Genetic variation in stomate densities and needle traits in a rangewide sampling of whitebark Pine (Pinus albicaulis)
- P. J. Vogan, Anna W. Schoettle. 2018. Associations between complete resistance to white pine blister rust and abiotic stress tolerances in limber pine (Pinus flexilis)
- Anna W. Schoettle, Richard A. Sniezko, John T. Kliejunas. 2018. Proceedings of the IUFRO joint conference: Genetics of five-needle pines, rusts of forest trees, and Strobusphere; 2014 June 15–20; Fort Collins, CO
- Christy M. Cleaver, Kelly S. Burns, Anna W. Schoettle. 2017. Limber Pine and White Pine Blister Rust Monitoring and Assessment Guide for Rocky Mountain National Park
- Mitra Menon, Justin C. Bagley, Christopher J. Friedline, Amy V. Whipple, Anna W. Schoettle, Alejandro Leal-Saenz, Christian Wehenkel, Francisco Molina-Freaner, Lluvia Flores-Renteria, M. Socorro Gonzalez-Elizondo, Richard A. Sniezko, Samuel A. Cushman, Kristen M. Waring, Andrew J. Eckert. 2017. The role of hybridization during ecological divergence of southwestern white pine (Pinus strobiformis) and limber pine (P. flexilis)
- Pam Allenstein, Jennifer DeWoody, David Gwaze, Valerie Hipkins, Gary Man, Anna W. Schoettle, Kirsty Shaw, Murphy Westwood. 2017. Improving genetic conservation of tree species
- Anna W. Schoettle, Jonathan C. Coop. 2017. Range-wide conservation of Pinus aristata: A genetic collection with ecological context for proactive management today and resources for tomorrow
- Sue Miller, Anna W. Schoettle, Kelly Burns, Richard Sniezko, Patricia A. Champ. 2017. Preempting the pathogen: Blister rust and proactive management of high-elevation pines
- Anna W. Schoettle. 2016. The growing knowledge base for limber pine - recent advances
- Patrick J. Vogan, Anna W. Schoettle. 2016. Carbon costs of constitutive and expressed resistance to a non-native pathogen in limber pine
- Jun-Jun Liu, Anna W. Schoettle, Richard A. Sniezko, Rona N. Sturrock, Arezoo Zamany, Holly Williams, Amanda Ha, Danelle Chan, Bob Danchok, Douglas P. Savin, Angelia Kegley. 2016. Genetic mapping of Pinus flexilis major gene (Cr4) for resistance to white pine blister rust using transcriptome-based SNP genotyping
- A. M. A. Casper, W. R. Jacobi, Anna W. Schoettle, K. S. Burns. 2016. Restoration planting options for limber pine (Pinus flexilis James) in the Southern Rocky Mountains
- Erin M. Borgman, Anna W. Schoettle, Amy L. Angert. 2015. Assessing the potential for maladaptation during active management of limber pine populations: A common garden study detects genetic differentiation in response to soil moisture in the Southern Rocky Mountains
- Anna W. Schoettle, Kelly S. Burns, J. Jeff Connor, Christy M. Cleaver. 2015. Limber pine conservation strategy: Recommendations for Rocky Mountain National Park
- Patrick J. Vogan, Anna W. Schoettle. 2015. Selection for resistance to white pine blister rust affects the abiotic stress tolerances of limber pine
- Erin M. Borgman, Anna W. Schoettle, Amy L. Angert. 2014. Using among-year variation to assess maternal effects in Pinus aristata and Pinus flexilis
- Anna W. Schoettle, Jeff Connor, John Mack, Phyllis Pineda Bovin, Jen Beck, Gretchen Baker, R. A. Sniezko, K. S. Burns. 2014. Establishing the science foundation to sustain high-elevation five-needle pine forests threatened by novel interacting stresses in four western National Parks [Proceedings]
- Anna W. Schoettle, R. A. Sniezko, A. Kegley, K. S. Burns. 2014. White pine blister rust resistance in limber pine: Evidence for a major gene
- Anna W. Schoettle, J. Connor, J. Mack, P. Pineda Bovin, J. Beck, G. M. Baker, R. A. Sniezko, K. S. Burns. 2013. Establishing the science foundation to sustain high-elevation five-needle pine forests threatened by novel interacting stresses in four western National Parks
- Jeff Connor, Anna W. Schoettle, Kelly Burns, Erin Borgman. 2012. Limber pine conservation in Rocky Mountain National Park
- Katarina Sweeney, Jeffrey Stone, Kathy Cook, Richard A. Sniezko, Angelia Kegley, Anna W. Schoettle. 2012. Needle reactions in resistance to Cronartium ribicola: Hypersensitivity response or not?
- Anna W. Schoettle, R. A. Sniezko, A. Kegley, R. Danchok, K. S. Burns. 2012. Patterns of resistance to Cronartium ribicola in Pinus aristata, Rocky Mountain bristlecone pine
- Kelly Burns, Jim Blodgett, Marcus Jackson, Brian Howell, William Jacobi, Anna W. Schoettle, Anne Marie Casper, Jennifer Klutsch. 2012. Monitoring limber pine health in the Rocky Mountains and North Dakota
- Anna W. Schoettle, J. G. Klutsch, R. A. Sniezko. 2012. Integrating regeneration, genetic resistance, and timing of intervention for the long-term sustainability of ecosystems challenged by non-native pests - a novel proactive approach
- Anna W. Schoettle, Betsy A. Goodrich, Valerie Hipkins, Christopher Richards, Julie Kray. 2012. Geographic patterns of genetic variation and population structure in Pinus aristata, Rocky Mountain bristlecone pine
- Robert E. Keane, D. F. Tomback, C. A. Aubry, A. D. Bower, E. M. Campbell, C. L. Cripps, M. B. Jenkins, M. F. Mahalovich, M. Manning, S. T. McKinney, M. P. Murray, D. L. Perkins, D. P. Reinhart, C. Ryan, Anna W. Schoettle, C. M. Smith. 2012. A range-wide restoration strategy for whitebark pine (Pinus albicaulis)
- S. G. Field, Anna W. Schoettle, J. G. Klutsch, S. J. Tavener, M. F. Antolin. 2012. Demographic projection of high-elevation white pines infected with white pine blister rust: a nonlinear disease model
- Anne Marie Casper, William R. Jacobi, Anna W. Schoettle, Kelly S. Burns. 2011. Restoration planting options for limber pines in Colorado and Wyoming
- Anne Marie Casper, William R. Jacobi, Anna W. Schoettle, Kelly S. Burns. 2011. Restoration planting options for limber pines in the southern Rocky Mountains
- Anna W. Schoettle, Michele Laskowski. 2011. High elevation white pines educational website
- Anna W. Schoettle, B. A. Goodrich, J. G. Klutsch, K. S. Burns, S. Costello, R. A. Sniezko. 2011. The proactive strategy for sustaining five-needle pine populations: An example of its implementation in the southern Rocky Mountains
- Jeffrey Stone, Anna W. Schoettle, Richard Sniezko, Angelia Kegley. 2011. Histological observations on needle colonization by Cronartium ribicola in susceptible and resistant seedlings of whitebark pine and limber pine
- Richard A. Sniezko, Mary F. Mahalovich, Anna W. Schoettle, Detlev R. Vogler. 2011. Past and current investigations of the genetic resistance to cronartium ribicola in high-elevation five-needle pines
- Anna W. Schoettle, Richard A. Sniezko, Angelia Kegley, Kelly S. Burns. 2011. Preliminary overview of the first extensive rust resistance screening tests of Pinus flexilis and Pinus aristata
- Sam Hendricks, Wendy Sutton, Jeffrey Stone, Richard Sniezko, Angelia Kegley, Anna W. Schoettle. 2011. Polymerase Chain Reaction (PCR) applications in white pine blister rust resistance screening
- Robert E. Keane, Anna W. Schoettle. 2011. Strategies, tools, and challenges for sustaining and restoring high elevation five-needle white pine forests in western North America
- Craig A. Bond, Patricia A. Champ, James Meldrum, Anna W. Schoettle. 2011. Investigating the optimality of proactive management of an invasive forest pest
- Jennifer G. Klutsch, Betsy A. Goodrich, Anna W. Schoettle. 2011. Limber pine forests on the leading edge of white pine blister rust distribution in Northern Colorado
- Anna W. Schoettle, Betsy A. Goodrich, Valerie Hipkins, Christopher Richards, Julie Kray. 2011. Geographic patterns of genetic variation, population structure and adaptive traits in Pinus aristata, Rocky Mountain bristlecone pine
- Elizabeth M. Campbell, Robert E. Keane, Evan R. Larson, Michael P. Murray, Anna W. Schoettle, Carmen Wong. 2011. Disturbance ecology of high-elevation five-needle pine ecosystems in western North America
- Jonathan D. Coop, Anna W. Schoettle. 2011. Fire and high-elevation, five-needle pine (Pinus aristata & P. flexilis) ecosystems in the southern Rocky Mountains: What do we know?
- Richard A. Sniezko, Anna W. Schoettle, Joan Dunlap, Detlev R. Vogler, David Conklin, Andrew Bower, Chris Jensen, Rob Mangold, Doug Daoust, Gary Man. 2011. Ex Situ gene conservation in high elevation white pine species in the United States-a beginning
- Anna W. Schoettle, J. G. Klutsch, M. F. Antolin, S. Field. 2011. A population genetic model for high-elevation five-needle pines: Projecting population outcomes in the presence of white pine blister rust
- Andrew D. Bower, Sierra C. McLane, Andrew Eckert, Stacy Jorgensen, Anna W. Schoettle, Sally Aitken. 2011. Conservation genetics of high elevation five-needle white pines
- Diana F. Tomback, Anna W. Schoettle, Mario J. Perez, Kristen M. Grompone, Sabine Mellmann-Brown. 2011. Regeneration and survival of whitebark pine after the 1988 Yellowstone fires
- Diana F. Tomback, Sheridan Samano, Elizabeth L. Pruett, Anna W. Schoettle. 2011. Seed dispersal in limber and southwestern white pine: Comparing core and peripheral populations
- Diana F. Tomback, Peter Achuff, Anna W. Schoettle, John W. Schwandt, Ron J. Mastrogiuseppe. 2011. The magnificent high-elevation five-needle white pines: Ecological roles and future outlook
- Kelly Burns, Jim Blodgett, Marcus Jackson, Brian Howell, William Jacobi, Anna W. Schoettle, Anne Marie Casper, Jennifer Klutsch. 2011. Monitoring limber pine health in the Rocky Mountains and North Dakota
- Anna W. Schoettle, Richard A. Sniezko, Angelia Kegley, Jerry Hill, Kelly S. Burns. 2010. Resistance to white pine blister rust in Pinus flexilis and P. aristata of the Southern Rockies - initial findings
- Craig A. Bond, Patricia A. Champ, James Meldrum, Anna W. Schoettle. 2010. Proactive or reactive? Optimal management of an invasive forest pest in a spatial framework
- Kelly Burns, Jim Blodgett, Dave Conklin, Brian W. Geils, Jim Hoffman, Marcus Jackson, William Jacobi, Holly Kearns, Anna W. Schoettle. 2010. White pine blister rust in the interior Mountain West
- Anne Marie Casper, William R. Jacobi, Anna W. Schoettle, Kelly S. Burns. 2010. Restoration planting options for limber pines impacted by mountain pine beetles and/or white pine blister rust in the Southern Rocky Mountains
- James T. Blodgett, Kelly S. Burns, Brian Howell, Marcus Jackson, William R. Jacobi, Anna W. Schoettle. 2010. Limber pine health survey in the Rocky Mountains and North Dakota
- Jonathan D. Coop, Robert T. Massatti, Anna W. Schoettle. 2010. Subalpine vegetation pattern three decades after stand-replacing fire: Effects of landscape context and topography on plant community composition, tree regeneration, and diversity
- Anna W. Schoettle, Richard A. Sniezko, Kelly S. Burns. 2009. Sustaining Pinus flexilis ecosystems of the southern Rocky Mountains (USA) in the presence of Cronartium ribicola and Dendroctonus ponderosae in a changing climate
- Jonathan D. Coop, Anna W. Schoettle. 2009. Regeneration of Rocky Mountain bristlecone pine (Pinus aristata) and limber pine (Pinus flexilis) three decades after stand-replacing fires
- Anna W. Schoettle, Kelly Burns, Sheryl Costello, Jeff Witcosky, Brian Howell, Jeff Connor. 2008. A race against beetles: Conservation of limber pine
- Richard A. Sniezko, Angelia Kegley, Robert Danchok, Anna W. Schoettle, Kelly S. Burns, Dave Conklin. 2008. Cronartium ribicola resistance in whitebark pine, southwestern white pine, limber pine and Rocky Mountain bristlecone pine - preliminary screening results from first tests at Dorena GRC
- Peter M. Brown, Anna W. Schoettle. 2008. Fire and stand history in two limber pine (Pinus flexilis) and Rocky Mountain bristlecone pine (Pinus aristata) stands in Colorado
- Kelly S. Burns, Anna W. Schoettle, William R. Jacobi, Mary Mahalovich. 2008. Options for the management of white pine blister rust in the Rocky Mountain Region
- Ann L. Lezberg, Michael A. Battaglia, Wayne D. Shepperd, Anna W. Schoettle. 2008. Decades-old silvicultural treatments influence surface wildfire severity and post-fire nitrogen availability in a ponderosa pine forest
- Anna W. Schoettle, Richard A. Sniezko, Kelly S. Burns, Freeman Floyd. 2007. Preparing the landscape for invasion - Early intervention approaches for threatened high elevation white pine ecosystems
- Anna W. Schoettle, Richard A. Sniezko. 2007. Proactive intervention to sustain high-elevation pine ecosystems threatened by white pine blister rust
- Anna W. Schoettle, K. S. Burns, F. Freeman, R. A. Sniezko. 2006. Threats, status & management options for bristlecone pines and limber pines in Southern Rockies
- Detlev R. Vogler, Annette Delfino-Mix, Anna W. Schoettle. 2006. White pine blister rust in high-elevation white pines: Screening for simply-inherited, hypersensitive resistance
- V. H. Bonnet, Anna W. Schoettle, W. D. Shepperd. 2005. Postfire environmental conditions influence the spatial pattern of regeneration for Pinus ponderosa
- Diana F. Tomback, Anna W. Schoettle, Kristen E. Chevalier, Cheri A. Jones. 2005. Life on the edge for limber pine: Seed dispersal within a peripheral population
- Anna W. Schoettle. 2004. Ecological roles of five-needle pine in Colorado: Potential consequences of their loss
- Anna W. Schoettle. 2004. Developing proactive management options to sustain bristlecone and limber pine ecosystems in the presence of a non-native pathogen
- V. H. Bonnet, Anna W. Schoettle, W. D. Shepperd. 2004. Spatial distribution of ponderosa pine seedlings along environmental gradients within burned areas in the Black Hills, South Dakota
- Pedro J. Aphalo, Anna W. Schoettle, Tarja Lehto. 2002. Leaf life span and the mobility of "non-mobile" mineral nutrients - the case of boron in conifers
- Andrew J. Schauer, Anna W. Schoettle, Richard L. Boyce. 2001. Partial cambial mortality in high-elevation Pinus aristata (Pinaceae)
- Anna W. Schoettle, Jose Negron. 2001. First report of two cone and seed insects on Pinus flexilis
- Michael F. Antolin, Anna W. Schoettle. 2001. Fragments, extinction, and recolonization: The genetics of metapopulations
- Jose Negron, Wayne A. Shepperd, Steve A. Mata, John B. Popp, Lance A. Asherin, Anna W. Schoettle, John M. Schmid, David A. Leatherman. 2001. Solar treatments for reducing survival of mountain pine beetle in infested ponderosa and lodgepole pine logs
- Anna W. Schoettle, S. G. Rochelle. 2000. Morphological variation of Pinus flexilis (Pinaceae), a bird-dispersed pine, across a range of elevations
- Anna W. Schoettle. 2000. Effect of two years of nitrogen deposition on shoot growth and phenology of Engelmann spruce seedlings
- W. H. Moir, Shannon G. Rochelle, Anna W. Schoettle. 1999. Microscale patterns of tree establishment near upper treeline, Snowy Range, Wyoming, USA
- Anna W. Schoettle, W. K. Smith. 1999. Interrelationships among light, photosynthesis and nitrogen in the crown of mature Pinus contorta ssp. latifolia
- Anna W. Schoettle, Kathy Tonnessen, John Turk, John Vimont, Robert Amundson, Ann Acheson, Janice Peterson. 1999. An assessment of the effects of human-caused air pollution on resources within the interior Columbia River basin.
- Anna W. Schoettle. 1998. Can we protect high-elevation wilderness vegetation from air pollution impacts?
- Anna W. Schoettle, William Moir. 1998. Terrestrial ecosystems [Chapter 4]
- Anna W. Schoettle. 1995. Predicting the potential sensitivity of high elevation wilderness vegetation to changes in atmospheric chemistry - a strategy
- Robert C. Musselman, D. G Fox, Anna W. Schoettle, C. M. Regan. 1994. Introduction [Chapter 1]
- Russell K. Monson, Michael C. Grant, Charles H. Jaeger, Anna W. Schoettle. 1992. Morphological causes for the retention of precipitation in the crowns of alpine plants
- W. K. Smith, Anna W. Schoettle, M. Cui. 1991. Importance of the method of leaf area measurement to the interpretation of gas exchange of complex shoots
- Douglas G. Fox, Anna W. Schoettle, Frank A. Vertucci. 1987. Glacier Lakes Ecosystem Experiment Site: an "Experimental" wilderness
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Program
https://research.fs.usda.gov/rmrs/programs/fweStation -
Project
https://research.fs.usda.gov/rmrs/projects/bil-whitebark-pine-resilience-networkStation -
Project
https://research.fs.usda.gov/rmrs/projects/bil-incorporating-adaptation-strategies-across-interior-west-adaptive-silvicultureStation -
Center or Group
https://research.fs.usda.gov/rmrs/centers/invasivespeciesworkinggroupStation
Multimedia
- Cornell University, Ithaca, NY, B.S., Biology - botany and biochemistry, 1978
- Cornell University, Ithaca, NY, M.S., Seed physiology and crop science, 1982
- University of Wyoming, Laramie, WY, Ph.D., Botany - ecophysiology and biophysics, 1990
Last updated July 7, 2026