Tamara Heartsill Scalley, PhD

Treesearch Publications

• Nafiseh Haghtalab, Tamara Heartsill Scalley, Tana E. Wood, J. Aaron Hogan, Humfredo Marcano-Vega, Thomas Brandeis, Thomas Ruzycki, Eileen Helmer. 2026. Multi-Resolution Mapping of Aboveground Biomass and Change in Puerto Rico’s Forests with Remote Sensing and Machine Learning
• Bianca M. Rodríguez-Cardona, Alonso Ramírez, Tamara Heartsill Scalley. 2026. Timing, magnitude, and sources of coarse particulate organic matter in tropical montane streams
• Katiana Garcia-Rosado, Ken M. Fritz, Augustin C. Engman, Tamara Heartsill Scalley, Sean P. Kelly, Carlos E. Ramos-Scharrón, Sofía Olivero-Lora. 2026. Foundational Elements for Developing a US Caribbean Streamflow Duration Assessment Method
• Mingjie Shi, Michael Keller, Barbara Bomfim, Lara Kueppers, Charlie Koven, Jessica Needham, Tamara Heartsill Scalley, L. Ruby. Leung. 2025. Assessing Simulations of Forest Hurricane Disturbance and Recovery in Puerto Rico by ELM‐FATES Using Field Measurements
• Sebastián Martinuzzi, Olga M. Ramos González, David P. Helmers, Tamara Heartsill Scalley, Volker C. Radeloff, Maya Quiñones. 2025. Defining and mapping the wildland-urban interface of Puerto Rico and the U.S. Virgin Islands:2010 Distribution and Implications
• Tania Lopez-Marrero, Tamara Heartsill Scalley. 2024. Tropical storms and ecological research in the Luquillo Experimental Forest: A bibliography
• Coral Matos, Amelia Merced, Tamara Heartsill Scalley. 2024. Habitat preferences and distribution of some common bryophytes in a tropical forest at the Luquillo Experimental Forest, Puerto Rico
• Virginia Hudspeth, Alberto Pastor, Tamara Heartsill Scalley. 2023. High-Light Demanding and Rare Tree Species Define Tropical Riparian Tree Communities
• Tamara Heartsill Scalley. 2023. Caribbean Islands (Springs of the World: Distribution, Ecology, and Conservation Status)
• Amelia Merced, Tamara Heartsill Scalley. 2022. Guía para las Briofitas de Puerto Rico
• Jiaying Zhang, Rafael L. Bras, Marcos Longo, Tamara Heartsill Scalley. 2022. Future Hurricanes Will Increase Palm Abundance and Decrease Aboveground Biomass in a Tropical Forest
• Sheng-I Yang, Thomas Brandeis, Eileen Helmer, Michael P. Oatham, Tamara Heartsill Scalley, Humfredo Marcano-Vega. 2022. Characterizing height-diameter relationships for Caribbean trees using mixed-effects random forest algorithm
• Jiaying Zhang, Rafael L. Bras, Marcos Longo, Tamara Heartsill Scalley. 2022. The impact of hurricane disturbances on a tropical forest: implementing a palm plant functional type and hurricane disturbance module in ED2-HuDi V1.0
• Jiaying Zhang, Tamara Heartsill Scalley, Rafael L. Bras. 2022. Parsing Long-Term Tree Recruitment, Growth, and Mortality to Identify Hurricane Effects on Structural and Compositional Change in a Tropical Forest
• Neftalí Ríos, Amelia Merced, Tamara Heartsill Scalley, Ariel Lugo. 2022. Acta CientíficaUna Revista Interdisciplinaria de Puerto Rico y el Caribe
• Neftali Rios, Amelia Merced, Tamara Heartsill Scalley, Ariel Lugo. 2022. Acta Científica. Edición Especial. Más allá de la resiliencia: Los efectos de los huracanes Irma y María en Puerto Rico y el Caribe.
• Jiaying Zhang, Tamara Heartsill Scalley, Rafael L. Bras. 2022. Forest Structure and Composition Are Critical to Hurricane Mortality
• William H. McDowell, Miguel C. Leon, Michelle D. Shattuck, Jody D. Potter, Tamara Heartsill Scalley, Grizelle Gonzalez, James B. Shanley, Adam S. Wymore. 2021. Luquillo Experimental Forest: Catchment science in the montane tropics
• Tamara Heartsill Scalley, Todd A. Crowl. 2021. Tropical forest understory riparian and upland composition, structure, and function in areas with different past land use
• ., Lawrence E. Stevens, Anwar A. Aly, Sarah M. Arpin, Iva Apostolova, Gail M. Ashley, Paulo Quadri Barba, Jose Barquín, Aude Beauger, Lahcen Benaabidate, Sami Ullah Bhat, Lhoussaine Bouchaou, Marco Cantonati, Teresa M. Carroll, Russell Death, Kathleen A. Dwire, Miguel Fernandes Felippe, Roderick J. Fensham, Alan E. Fryar, Roger Pascual i Garsaball, Vojsava Gjoni, Douglas S. Glazier, Nico Goldscheider, Joseph T. Gurrieri, Ragnhildur Guðmundsdóttir, Atzalan Rodriguez Guzman, Michal Hájek, Kristian Hassel, Tamara Heartsill Scalley, Jaume Solé i Herce, Dirk Hinterlang, Joseph H. Holway, Jari Ilmonen, Jeffrey Jenness, Jutta Kapfer, Ioannis Karaouzas, Robert L. Knight, Agnes-Katharina Kreiling, Christian Herrera Lameli, Jeri D. Ledbetter, Nataly Levine, Melinda D. Lyons, Robert E. Mace, Angeliki Mentzafou, Pierre Marle, Nils Moosdorf, Monica K. Norton, Allan Pentecost, Guillermo García Pérez, Bianca Perla, Abdullah A. Saber, Donald Sada, Stefano Segadelli, Kamilla Skaalsveen, Abraham E. Springer, Susan K. Swanson, Benjamin F. Schwartz, Peter Sprouse, Memory Tekere, Benjamin W. Tobin, Ernest Azwindini Tshibalo, Olivier Voldoire. 2021. The Ecological Integrity of Spring Ecosystems: A Global Review
• N.M. Soto-Parrilla, Tamara Heartsill Scalley. 2021. Missing the Trees for the Forest: Post-Hurricane Understory Vegetation in Relation to Spatial Variation
• M.C. Leon, Tamara Heartsill Scalley, I. Santiago, W. H. McDowell. 2021. Hydrological Mapping in the Luquillo Experimental Forest: New Local Datum Improves Watershed Ecological Knowledge
• Ariel Lugo, Oscar J. A. Martínez, Ernesto Medina, Gerardo Aymard, Tamara Heartsill Scalley. 2020. Novelty in the tropical forests of the 21st century
• Tamara Heartsill Scalley, Gerónimo Mercado. 2019. Sound Circles/Inmersion
• Tania López-Marrero, Tamara Heartsill Scalley, Carlos F. Rivera-López, Isabel A. Escalera-García, Mariangelí Echevarría-Ramos. 2019. Broadening Our Understanding of Hurricanes and Forests on the Caribbean Island of Puerto Rico: Where and What Should We Study Now?
• M. R. Willig, L. Woolbright, S. J. Presley, T. D. Schowalter, R. B. Waide, Tamara Heartsill Scalley, J. K. Zimmerman, Grizelle Gonzalez, Ariel Lugo. 2019. Populations are not declining and food webs are not collapsing at the Luquillo Experimental Forest
• Mei Yu, Elsie Rivera-Ocasio, Tamara Heartsill Scalley, Daniel Davila-Casanova, Neftalí Rios-López, Qiong Gao. 2019. Landscape-Level Consequences of Rising Sea-Level on Coastal Wetlands: Saltwater Intrusion Drives Displacement and Mortality in the Twenty-First Century
• John Rothlisberger, Tamara Heartsill Scalley, Russ F. Thurow. 2017. The role of wild and scenic rivers in the conservation of aquatic biodiversity
• Tamara Heartsill Scalley. 2017. Insights on Forest Structure and Composition from Long-Term Research in the Luquillo Mountains
• H. Manrique-Hernandez, Tamara Heartsill Scalley, M. Barreto-Orta, C. M. Betancourt-Roman, J. R. Ortiz-Zayas. 2016. Assessing restoration outcomes in light of succession: management implications for tropical riparian forest restoration
• Tamara Heartsill Scalley, Saara DeWalt, François Korysko, Guy Van Laere, Kasey Jacobs, Seth Panka, Joseph Torres. 2016. Communication from the Information Sharing Working Group: Agreement for Data Sharing Among Caribbean Foresters
• Tamara Heartsill Scalley, Grizelle Gonzalez. 2016. Introduction: Caribbean forest dynamics and regional forestry initiatives
• Grizelle Gonzalez, Tamara Heartsill Scalley. 2016. Building a collaborative network to understand regional forest dynamics and advance forestry initiatives in the Caribbean
• Tania del M. Lopez-Marrero, Tamara Heartsill Scalley. 2014. Land-cover composition, water resources and land management in the watersheds of the Luquillo Mountains, northeastern Puerto Rico.
• M. Pfeifer, Tamara Heartsill Scalley. 2014. BIOFRAG - a new database for analyzing BIOdiversity responses to forest FRAGmentation
• Ariel Lugo, Tamara Heartsill Scalley. 2014. Research in the Luquillo Experimental Forest has advanced understanding of tropical forests and resolved management issues
• G. E. Ponce Campos, M. S. Moran, A. Huete, Y. Zhang, C. Bresloff, T.E. Huxman, D. Eamus, D. D. Bosch, A. R. Buda, S. A. Gunter, Tamara Heartsill Scalley, S. G. Kitchen, M. P. McClaran, Henry McNab, Diane Sutherland, J. A. Morgan, D. P. C. Peters, E. J. Sadler, M. S. Seyfried, P. J. Starks. 2013. Ecosystem resilience despite large-scale altered hydroclimatic conditions
• Tamara Heartsill Scalley. 2012. Vegetation
• James Vose, Chelcy R. Ford, Stephanie H. Laseter, Salli Dymond, Ge Sun, Mary Beth Adams, Stephen D. Sebestyen, John Campbell, Charlie H. Luce, Devendra M. Amatya, Kelly J. Elder, Tamara Heartsill Scalley. 2012. Can forest watershed management mitigate climate change impacts on water resources?
• E. López Machado, K. Soto Hidalgo, Tamara Heartsill Scalley. 2012. Composición y estructura de un bosque tropical urbano en el karso norteno de Puerto Rico
• T. López-Marrero, Tamara Heartsill Scalley. 2012. Get up, stand up:environmental situation, threats, and opportunities in the insular Caribbean
• T. M. López Marrero, K. Yamane, Tamara Heartsill Scalley, N. Villanueva Colón. 2012. Cartographic essay: the various shapes of the insular Caribbean: population and environment
• Tamara Heartsill Scalley. 2012. Freshwater resources in the insular Caribbean: an environmental perspective
• Tamara Heartsill Scalley, F.N. Scatena, Samuel Moya, Ariel Lugo. 2012. Long-term dynamics of organic matter and elements exported as coarse particulates from two Caribbean montane watersheds
• James Vose, Chelcy F. Miniat, Stephanie H. Laseter, Salli Dymond, Ge Sun, Mary Beth Adams, Stephen D. Sebestyen, John Campbell, Charlie H. Luce, Devendra M. Amatya, Kelly J. Elder, Tamara Heartsill Scalley. 2012. Can forest watershed management mitigate climate change effects on water resources
• Nancy L. Harris, Ariel Lugo, Sandra Brown, Tamara Heartsill Scalley. 2012. Luquillo Experimental Forest: Research History and Opportunities
• Alejandro A. Royo, Tamara Heartsill Scalley, Samuel Moya, Fred N. Scatena. 2011. Non-arborescent vegetation trajectories following repeated hurricane disturbance: ephemeral versus enduring responses
• Tamara Heartsill Scalley, Frederick N. Scatena, Ariel Lugo, Samuel Moya, Carlos R. Estrada Ruiz. 2010. Changes in structure, composition, and nutrients during 15 years of hurricane-induced succession in a subtropical wet forest in Puerto Rico
• Tamara Heartsill Scalley, Todd A. Crowl, Jill Thompson. 2009. Tree species distributions in relation to stream distance in mid-montane wet forest, Puerto Rico
• Rachel Pérez, Tamara Heartsill Scalley. 2008. Root nodulation in the wetland tree Pterocarpus officinalis along coastal and montane systems of Northeast of Puerto Rico
• Tamara Heartsill Scalley, F.N. Scatena, Carlos R. Estrada Ruiz, W.H. McDowell, Ariel Lugo. 2007. Disturbance and long-term patterns of rainfall and throughfall nutrient fluxes in a subtropical wet forest in Puerto Rico
• Todd A. Crowl, Vanessa Welsh, Tamara Heartsill Scalley. 2006. Effects of different types of conditioning on rates of leaf-litter shredding by Xiphocaris elongata, a Neotropical freshwater shrimp.
• Tamara Heartsill Scalley, Carlos R. Estrada Ruiz, Samuel Moya. 2006. Patrones de lluvia, transcolación y flujo de nutrientes en las cuencas de Bisley, Bosque Experimental de Luquillo, Puerto Rico
• Alan P. Covich, Todd A. Crowl, Tamara Heartsill Scalley. 2006. Effects of drought and hurricane disturbances on headwater distributions of palaemonid river shrimp (Macrobrachium spp.) in the Luquillo Mountains, Puerto Rico.

Data and Tools

• Dataset
  

  Silvics of North America

  https://research.fs.usda.gov/products/dataandtools/silvics-north-america
• Dataset
  

  Tree census at Bisley Experimental Watersheds three months after Hurricane Maria

  https://research.fs.usda.gov/iitf/products/dataandtools/tree-census-bisley-experimental-watersheds-three-months-after-hurricane

• Project
  

  Caribbean Foresters

  https://research.fs.usda.gov/iitf/projects/caribbean-foresters
  Station

  International Institute of Tropical Forestry
• Project
  

  Forest's Response to Hurricanes

  https://research.fs.usda.gov/iitf/projects/forest-responses-hurricanes
  Station

  International Institute of Tropical Forestry

Other Publications

• User Guide
  

  Bryophytes of El Yunque National Forest, Puerto Rico (Quick reference guide)

  https://research.fs.usda.gov/iitf/understory/bryophytes-el-yunque-national-forest-puerto-rico-quick-reference-guide
• User Guide
  

  Bryophytes of Urban and Community Forests, Puerto Rico (Quick reference guide)

  https://research.fs.usda.gov/iitf/understory/bryophytes-urban-and-community-forests-puerto-rico-quick-reference-guide
• Fact Sheet
  

  Sphagnum portoricense, El Yunque National Forest

  https://research.fs.usda.gov/iitf/understory/sphagnum-portoricense-el-yunque-national-forest
• User Guide
  

  Sphagnum species in Puerto Rico (Quick reference guide)

  https://research.fs.usda.gov/iitf/understory/sphagnum-species-puerto-rico-quick-reference-guide

Multimedia

• Webinars
  

  Identificación de briofitas en Puerto Rico

  https://research.fs.usda.gov/iitf/products/multimedia/webinars/identificacion-de-briofitas-en-puerto-rico
• Video

  Tamara Heartstill-Scalley, Research Ecologist

  https://research.fs.usda.gov/iitf/products/multimedia/videos/tamara-heartstill-scalley-research-ecologist

• Utah State University, Logan, Ph.D., Ecology, 2005
• University of Puerto Rico, Río Piedras, M.Sc., Biology, 2000
• University of Puerto Rico, Río Piedras, B.A., Geography, 1994