Luquillo Experimental Forest
Located in Puerto Rico, the 11,540 ha (28,516 acres) Luquillo Experimental Forest is the largest tropical forest in the experimental forest and range system and the only experimental forest with the same boundary as the national forest, that is, El Yunque National Forest and the Luquillo are one. It has been studied for over 100 years and is one of the most intensively studied tropical forests in the world. The Luquillo Experimental Forest comprises the largest area of primary forests in Puerto Rico and the most pristine rivers in the island, including several wild and scenic rivers.

History 1876-2017
- 1876: King of Spain Alphonso XII proclaims 10 000 ha (24,710 acres) of forested land in the Luquillo Mountains as a forest reserve.
- 1903: President Theodore Roosevelt establishes the Luquillo Forest Reserve.
- 1956: The forest is designated the Luquillo Experimental Forest to recognize the growing importance of research.
- 1976: The forest is designated as a biosphere reserve by the United Nations.
- 1988: The Long Term Ecological Research (LTER) program begins in collaboration with the University of Puerto Rico.
- 2005: El Toro Wilderness Area is established.
- 2009: The Luquillo Critical Zone Observatory (LCZO) is established within El Yunque.
- 2017: A new management plan for El Yunque National Forest in final revision process.
Climate
Located within the Holdridge Life Zone Classification System, the Luquillo Experimental Forest is a subtropical moist to subtropical rain forest. Along the 1,000-meter elevation gradient (100 to 1,075 m asl), mean annual temperature ranges from 25 to 19 °C and annual rainfall from 2,000 to 5,000 mm. The average annual rainfall for the forest is 3,880 mm. The climate is subtropical maritime andis moderated by steady trade winds, which maintainrelatively constant year-round air temperatures. The annual temperature variation within a climate station is only 3.0 to 3.5 °C. Rainfall intensity varies at all temporal scales from diurnal to decadal. Although all months receive in excess of 100 mm, there is a period of lower rainfall between February and April and peaks of rainfall in September. Storms and hurricanes are climatic events that dramatically change forest conditions. For example, winds normally range from 8 to 18 km/h but exceed 200 km/h during category 5 hurricanes. Rainfall can exceed 400 mm/day during storms and low-pressure systems. Such events trigger numerous landslides. Droughts (< 100 mm/month) recur on decadal scales, but natural fires do not occur.
There are four soil associations representing 19 soil series. The principal orders are Ultisols and Inceptisols, which respectively occupy 50 and 20 percent of the total forest area, and small areas with Inceptisols and other soil types. Dominant soils are deep, highly weathered and leached clays, with low pH and base saturation less than 35 percent at a depth of 1.25 m.
Databases
Numerous data bases are compiled at the Luquillo Experimental Forest, including maps of geology, soils, vegetation, and disturbance history; ten sets of aerial photography (since 1936); air temperature and precipitation (since 1909); solar radiation, windspeed and direction, relative humidity; NADP precipitation (since 1984); as well as chemical composition of precipitation, dryfall,cloudwater, bulk soil, soil solution, throughfall, stemflow, streamwater, plant, animal, and fungal tissue; vegetative composition, above- and below-ground biomass, tree growth/mortality, litterfall, litter decomposition, wood decomposition, mycorrhizal associations, phenology; and population records and biomass of terrestrial and aquatic fauna. Many long-term data records are available on the USFS Research Data Archive and the Luquillo LTER web page. [External Site: Opens in New Window]
Research and Opportunities
Research studies in the Luquillo Experimental Forest range from plantation forestry, reforestation, land rehabilitation, tree growth, and ecosystem structure and function, to wildlife ecology, hydrology, geomorphology, and disturbance and landscape ecology. There are more than 30 cross-site experiments, including:
- LIDET (Long-Term Intersite Decomposition Experiment Team)
- LINX (Lotic Intersite Nitrogen Experiment)
- DONIC (Dissolved Organic Nitrogen Intersite Comparison)
- WW-DECOEX (World Wide Aquatic Leaf Decomposition Experiment)
- Earthworms and soil processes in tropical ecosystems
- Relationship between nutrient inputs and faunal diversity
- Canopy herbivory and soil processes in a temperate and tropical forest
- Global forest dynamics network
- Landscape fragmentation and forest fuel accumulation; effects of fragment size, age, and climate.
The Luquillo Experimental Forest offers opportunities for research in tropical forestry and related fields. Programs such as those of the USDA Forest Service, the National Science Foundation, the USDI Geological Survey, and the University of Puerto Rico are available for research collaboration in all fields of the natural sciences.
Facilities Information
The Sabana Field Research Station
The Sabana Field Research Station is a facility of the USDA Forest Service’s International Institute of Tropical Forestry (Institute), a center for excellence where creativity and accomplishments result in timely products and services that anticipate the needs of society as it mitigates and adapts to environmental change.
Our mission is to develop and disseminate scientifically-based knowledge that contributes to the conservation of forests, wildlife, and watersheds of the American tropics in the context of environmental change. Furthermore, the Institute strives to cultivate an inclusive environment and ensure that all who visit our facilities are provided equal access to opportunities, programs, and services that are available.
The El Verde Field Station

The El Verde Field Station, which has living quarters and laboratory facilities for up to 25 scientists, is maintained by the Institute for Tropical Ecosystem Studies in the Luquillo. The field station provides collections of local plants and animals; a laboratory with light meters, balances, microscopes, pH meters, hoods, etc.; line power backed up by a generator; and gas, air, and vacuum lines. The Biology Department of the University of Puerto Rico (Río Piedras) maintains a second field station in the palo colorado forest of the Luquillo Experimental Forest.
Among the resources available through the Institute are the Bisley Experimental Watersheds, the Sabana Field Research Station, a woodshop, analytical laboratory for soils and vegetation, tropical forestry library with 55,000 documents, 10,000 bound volumes, 100 journal subscriptions, map, film, and slide collections of the entire Oxford forestry collection on microfilm, FAO documents and journal listings from larger libraries (e.g., Oxford, University of Georgia, University of Florida, USDA National Agricultural Library) in microfiche, computerized literature searching facilities, and a research herbarium with more than 95 percent of the 700+ tree species in Puerto Rico. The herbarium is operated by the Botanical Garden of the University of Puerto Rico.
Contact
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Long-term Monitoring and Data
The Bisley Experimental Watersheds (BEW) are three adjacent watersheds managed by the Institute, located in tabonuco forest a few kilometers south of the Sabana Field Research Station.
Within the BEW, a number of long-term ecological, hydrological, and silvicultural research projects have been conducted since the 1980s, carried out by Institute scientists and collaborating researchers from the Luquillo Long-Term Ecological Research Program (LUQ-LTER), the Luquillo Critical Zone Observatory (LCZO), and the U.S. Geological Survey, among others.

Long-term, detailed monitoring of biogeochemistry, climate variables, and stream runoff in Bisley has produced some of the longest environmental records on watershed processes in the Neotropics, and has made significant contributions to the understanding of hydrologic, carbon, nitrogen, and phosphorus cycling in tropical wet forests.
The data from Bisley provide an unequalled opportunity to understand seasonal and inter-annual climatic variability and quantify its relationships with ecosystem processes, including the effects of multiple hurricanes, droughts, plant aboveground stoichiometry, and forest diversity and structure on biogeochemistry and faunal assemblages at the plot and watershed scales. They also contribute to collaborative analysis and research with ecologists from Coweeta Hydrologic Laboratories and other experimental forests and watersheds to support synthesis and large-scale understanding of climate change parameters and stream discharge relationships.
Bisley Facilities

Research facilities include two walk-up canopy towers, climate stations, three stream gages, throughfall and litter collectors, permanent ecological study plots, and areas designated for aquatic research. Forest composition and soil and plant nutrient content are monitored in 83 permanent 10 meter diameter plots on a 40 x 40 meter grid every 5 years.
Canopy throughfall and litterfall are measured weekly. Stream chemistry and coarse particulate export are measured every two weeks in three streams. Shrimp abundances are measured twice yearly in two BEW streams, and algal abundance, ecosystem metabolism, chlorophyll a, and insect abundance are measured biannually in one BEW stream.
Treesearch Publications
- Kai L. Kopecky, Max C. N. Castorani, Ty Tuff, Audrey Barker Plotkin, David M. Bell, Jill F. Johnstone, John S. Kominoski, Jesse B. Nippert, Christopher J. Nytch, Steven C. Pennings, Daniel C. Reed, Aaron B. Shiels, Katharine N. Suding. 2026. Legacies of foundation species shape life after death
- Omar Gutierez del Arroyo-Santiago, Hartman Melannie D., Silver Whendee. 2025. Modeling the Effects of Increased Hurricane Frequency on the Tropical Forest Carbon Cycle
- Tana E. Wood, Tucker Colin, Aura M. Alonso-Rodríguez, M. Isabel Loza, Iana F. Grullón-Penkova, Molly A. Cavaleri, Sasha C. Reed. 2025. Warming induces unexpectedly high soil respiration in a wet tropical forest
- H. E. Erickson, Grizelle Gonzalez, W.H. McDowell, J.D. Potter. 2025. Increasingly conservative N cycling in a wet tropical forest: Litter and stream N concentrations decline over 29 years despite surges from hurricanes
- Stephen D. Sebestyen, James B. Shanley, Theresa Blume, Jonathan M. Duncan, Julia Jones, Catalina Segura, M. Alisa. Mast. 2025. Introduction to the Special Issue on Research and Observatory Catchments
- Mingjie Shi, Michael Keller, Barbara Bomfim, Lara Kueppers, Charlie Koven, Jessica Needham, Tamara Heartsill Scalley, L. Ruby. Leung. 2025. Assessing Simulations of Forest Hurricane Disturbance and Recovery in Puerto Rico by ELM‐FATES Using Field Measurements
- Yanchen Sun, Yongchao Yin, Guang He, Gyuhyon Cha, Héctor Ayala-del-Río, Grizelle Gonzalez, Konstantinos Konstantinidis, Frank E. Löffler. 2024. pH selects for distinct N2O-reducing microbiomes in tropical soil microcosms
- Tania Lopez-Marrero, Tamara Heartsill Scalley. 2024. Tropical storms and ecological research in the Luquillo Experimental Forest: A bibliography
- Coral Matos, Amelia Merced, Tamara Heartsill Scalley. 2024. Habitat preferences and distribution of some common bryophytes in a tropical forest at the Luquillo Experimental Forest, Puerto Rico
- Mingjie Shi, Michael Keller, Barbara Bomfim, Lingcheng Li, Charlie Koven, Lara Kueppers, Ryan Knox, Jessica Needham, Shih‐Chieh Kao, Peter E. Thornton, Michele M. Thornton, L. Ruby. Leung. 2024. Functionally Assembled Terrestrial Ecosystem Simulator (FATES) for Hurricane Disturbance and Recovery
- Joel A. Mercado-Díaz, Eva Holupchinski, Nora Alvarez-Berrios, William A. Gould, Paul Miller, Thomas Mote, Craig Ramseyer, Grizelle Gonzalez. 2023. Fostering Knowledge Exchange and Collaboration among Drought-Related Initiatives in the Caribbean
- Rob Tunison, Tana E. Wood, Sasha C. Reed, Molly A. Cavaleri. 2023. Respiratory Acclimation of Tropical Forest Roots in Response to In Situ Experimental Warming and Hurricane Disturbance
- Virginia Hudspeth, Alberto Pastor, Tamara Heartsill Scalley. 2023. High-Light Demanding and Rare Tree Species Define Tropical Riparian Tree Communities
- Alberto C. Cruz-Mendoza, Frank F. Rivera-Milan, Wayne J. Arendt, Laura L. Fidalgo-De Souza, Jessica Ilse, Joseph M. Wunderle Jr. 2023. Effects of Hurricanes Irma and Maria on Abundance and Occupancy of Margarops fuscatus (Pearly-eyed Thrasher) in the Luquillo Experimental Forest, Puerto Rico
- Dingfang Chen, Mei Yu, Grizelle Gonzalez, Qiong Gao. 2023. Altitudinal Pattern of Soil Organic Carbon and Nutrients in a Tropical Forest in Puerto Rico
- Eileen Helmer, Shannon Kay, Humfredo Marcano-Vega, Jennifer S. Powers, Tana E. Wood, Xiaolin Zhu, David Gwenzi, Thomas S. Ruzycki. 2023. Multiscale predictors of small tree survival across a heterogeneous tropical landscape
- Ashley Van Beusekom, Grizelle Gonzalez, María M. Rivera. 2023. Tropical Forest Microclimatic Changes: Hurricane, Drought, and 15–20 Year Climate Trend Effects on Elevational Gradient Temperature and Moisture
- Ariel Lugo, Mary Jean Sanchez, Luz S. Betancourt, Olga Ramos, Ivan Vicens. 2023. A Stoichiometric Comparison of Primary and Secondary Forest Stands on Acid Soils
- Amelia Merced, Tamara Heartsill Scalley. 2022. Guía para las Briofitas de Puerto Rico
- J. Aaron Hogan, Joanne M. Sharpe, Ashley Van Beusekom, Sarah Stankavich, Samuel Matta Carmona, John E. Bithorn, Jamarys Torres-Díaz, Grizelle Gonzalez, Jess K. Zimmerman, Aaron B. Shiels. 2022. Solar radiation and soil moisture drive tropical forest understory responses to experimental and natural hurricanes
- Ivia I Moreno, Maria F. Barberena‐Arias, Grizelle Gonzalez, D. Jean Lodge, Sharon A. Cantrell. 2022. Canopy opening increases leaf‐shredding arthropods and nutrient mineralization but not mass loss in wet tropical forest
- D. Jean Lodge, Ashley Van Beusekom, Grizelle Gonzalez, Mareli Sánchez‐Julia, Sarah Stankavich. 2022. Disturbance reduces fungal white‐rot litter mat cover in a wet subtropical forest
- Ariel Lugo, Neftali Rios. 2022. Acta Cientifica: Edicion Especial. Frank Howard Wadsworth(1915-2022): un tributo al cientifico, mentor, y amigo.
- Jiaying Zhang, Rafael L. Bras, Marcos Longo, Tamara Heartsill Scalley. 2022. Future Hurricanes Will Increase Palm Abundance and Decrease Aboveground Biomass in a Tropical Forest
- Georg Wiehl, Christian Wirth, Michaela J. Woods, Paul-Camilo Zalamea, Amy E. Zanne, Habacuc Flores-Moreno, Jeff R. Powell, William K. Cornwell, James W. Dalling, Amy T. Austin, Aimée T. Classen, Paul Eggleton, Kei-ichi Okada, Catherine L. Parr, E. Carol Adair, Stephen Adu-Bredu, Md Azharul Alam, Carolina Alvarez-Garzón, Deborah Apgaua, Roxana Aragón, Marcelo Ardon, Stefan K. Arndt, Louise A. Ashton, Nicholas A. Barber, Jacques Beauchêne, Matty P. Berg, Jason Beringer, Matthias M. Boer, José Antonio Bonet, Katherine Bunney, Tynan J. Burkhardt, Dulcinéia Carvalho, Dennis Castillo-Figueroa, Lucas A. Cernusak, Alexander W. Cheesman, Tainá M. Cirne-Silva, Jamie R. Cleverly, Johannes H. C. Cornelissen, Timothy J. Curran, André M. D’Angioli, Caroline Dallstream, Nico Eisenhauer, Fidele Evouna Ondo, Alex Fajardo, Romina D. Fernandez, Astrid Ferrer, Marco A. L. Fontes, Mark L. Galatowitsch, Grizelle Gonzalez, Felix Gottschall, Peter R. Grace, Elena Granda, Hannah M. Griffiths, Mariana Guerra Lara, Motohiro Hasegawa, Mariet M. Hefting, Nina Hinko-Najera, Lindsay B. Hutley, Jennifer R. Jones, Anja Kahl, Mirko Karan, Joost A. Keuskamp, Tim Lardner, Michael Liddell, Craig Macfarlane, Cate Macinnis-Ng, Ravi F. Mariano, M. Soledad Méndez, Wayne S. Meyer, Akira S. Mori, Aloysio S. Moura, Matthew Northwood, Romà Ogaya, Rafael S. Oliveira, Alberto Orgiazzi, Juliana Pardo, Guille Peguero, Josep Penuelas, Luis I Perez, Juan M. Posada, Cecilia M. Prada, Tomáš Přívětivý, Suzanne M. Prober, Jonathan Prunier, Gabriel W. Quansah, Víctor Resco de Dios, Ronny Richter, Mark P. Robertson, Lucas F. Rocha, Megan A. Rúa, Carolina Sarmiento, Richard P. Silberstein, Mateus C. Silva, Flávia Freire Siqueira, Matthew Glenn Stillwagon, Jacqui Stol, Melanie K. Taylor, François P. Teste, David Y. P. Tng, David Tucker, Manfred Türke, Michael Ulyshen, Oscar J. Valverde-Barrantes, Eduardo van den Berg, Richard S. P. van Logtestijn, G. F. (Ciska) Veen, Jason G. Vogel, Timothy J. Wardlaw. 2022. Termite sensitivity to temperature affects global wood decay rates
- Ashley Van Beusekom, Grizelle Gonzalez. 2022. Near-Surface Atmospheric Behavior over Complex Tropical Topography in Puerto Rico Dominated by Seasonal Patterns Despite Frequent Environmental Changes
- Aura M. Alonso-Rodríguez, Tana E. Wood, Jamarys Torres-Díaz, Molly A. Cavaleri, Sasha C. Reed, Benedicte Bachelot. 2022. Understory plant communities show resistance to drought, hurricanes, and experimental warming in a wet tropical forest
- Jiaying Zhang, Rafael L. Bras, Marcos Longo, Tamara Heartsill Scalley. 2022. The impact of hurricane disturbances on a tropical forest: implementing a palm plant functional type and hurricane disturbance module in ED2-HuDi V1.0
- K. E. Clark, R. F. Stallard, S. F. Murphy, M. A. Scholl, Grizelle Gonzalez, A. F. Plante, W. H. McDowell. 2022. Extreme rainstorms drive exceptional organic carbon export from forested humid-tropical rivers in Puerto Rico
- Wei Huang, Grizelle Gonzalez, María Fernanda Barberena-Arias, Xiaoming Zou. 2022. Predominant climate control on green leaf decomposition across subtropical rain and dry forests following hurricane disturbances
- Jiaying Zhang, Tamara Heartsill Scalley, Rafael L. Bras. 2022. Parsing Long-Term Tree Recruitment, Growth, and Mortality to Identify Hurricane Effects on Structural and Compositional Change in a Tropical Forest
- Neftalí Ríos, Amelia Merced, Tamara Heartsill Scalley, Ariel Lugo. 2022. Acta CientíficaUna Revista Interdisciplinaria de Puerto Rico y el Caribe
- Neftali Rios, Amelia Merced, Tamara Heartsill Scalley, Ariel Lugo. 2022. Acta Científica. Edición Especial. Más allá de la resiliencia: Los efectos de los huracanes Irma y María en Puerto Rico y el Caribe.
- Jiaying Zhang, Tamara Heartsill Scalley, Rafael L. Bras. 2022. Forest Structure and Composition Are Critical to Hurricane Mortality
- William H. McDowell, Miguel C. Leon, Michelle D. Shattuck, Jody D. Potter, Tamara Heartsill Scalley, Grizelle Gonzalez, James B. Shanley, Adam S. Wymore. 2021. Luquillo Experimental Forest: Catchment science in the montane tropics
- Ariel E. Lugo. 2021. The Luquillo Experimental Forest: A Neotropical example of the interaction between forest conservation and long-term ecological research
- Julie C. Zinnert, Jesse B. Nippert, Jennifer A. Rudgers, Steven C. Pennings, Grizelle Gonzalez, Merryl Alber, Sara G. Baer, John M. Blair, Adrian Burd, Scott L. Collins, Christopher Craft, Daniela Di Iorio, Walter K. Dodds, Peter M. Groffman, Ellen Herbert, Christine Hladik, Fan Li, Marcy E. Litvak, Seth Newsome, John O’Donnell, William T. Pockman, John Schalles, Donald R. Young. 2021. State changes: insights from the U.S. Long Term Ecological Research Network
- Tamara Heartsill Scalley, Todd A. Crowl. 2021. Tropical forest understory riparian and upland composition, structure, and function in areas with different past land use
- Mariela Encarnación Ojeda, Elvia J. Meléndez Ackerman, Rayza Hernández Muñiz. 2021. Environmental correlates of leaf stress symptoms in the micro-orchid Lepanthes woodburyana following Hurricane María
- Rayza M. Hernández Muñiz, Elvia J. Meléndez Ackerman, Raymond L. Tremblay, Omar A. Monsegur Rivera. 2021. Short-term effects of Hurricane María on populations size and individual survival in Lepanthes eltoroensis (Orchidaceae) at the Luquillo Experimental Forest, Puerto Rico
- T.J. Hawley Matlaga, P.A. Burrowes, R. Hernández-Pacheco, J. Pena, C. Sutherland, Tana E. Wood. 2021. Warming increases activity in the common tropical frog Eleutherodactylus coqui
- Xiaolin Zhu, Eileen Helmer, David Gwenzi, Melissa Collin, Sean Fleming, Jiaqi Tian, Humfredo Marcano-Vega, Elvia J. Meléndez-Ackerman, Jess K. Zimmerman. 2021. Characterization of Dry-Season Phenology in Tropical Forests by Reconstructing Cloud-Free Landsat Time Series
- TaeOh Kwon, Hideaki Shibata, Sebastian Kepfer-Rojas, Inger K. Schmidt, Klaus S. Larsen, Claus Beier, Björn Berg, Kris Verheyen, Jean-Francois Lamarque, Frank Hagedorn, Nico Eisenhauer, Ika Djukic, Grizelle Gonzalez, Ian Yesilonis, Lindsey Rustad. 2021. Effects of Climate and Atmospheric Nitrogen Deposition on Early to Mid-Term Stage Litter Decomposition Across Biomes
- Marconi Campos-Cerqueira, Adam J. Terando, Brent A. Murray, Jaime A. Collazo, T. Mitchell. Aide. 2021. Climate change is creating a mismatch between protected areas and suitable habitats for frogs and birds in Puerto Rico
- Daniela Yaffar, Tana E. Wood, Sasha C. Reed, Benjamin L. Branoff, Molly A. Cavaleri, Richard J. Norby. 2021. Experimental warming and its legacy effects on root dynamics following two hurricane disturbances in a wet tropical forest
- Kelsey R. Carter, Tana E. Wood, Sasha C. Reed, Kaylie M. Butts, Molly A. Cavaleri. 2021. Experimental warming across a tropical forest canopy height gradient reveals minimal photosynthetic and respiratory acclimation
- Claudia M. Ospina-Sanchez, Jose G. Palacios-Vargas, Grizelle Gonzalez. 2021. A new species of Pronura (Neanuridae: Paleonurini) from the Luquillo Mountains, Puerto Rico with a key to American poorly tuberculated Paleonurini
- Elvis Torres-Delgado, Grizelle Gonzalez, Ernesto Medina, María M. Rivera. 2021. A Multiyear Record of Rainfall and Ionic Composition along an Elevation Gradient in Northeastern Puerto Rico
- Sandy M. Smith, Michael Steinwandter, Katalin Szlavecz, José Antonio Talavera, Dolores Trigo, Jiro Tsukamoto, Sheila Uribe-López, Anne W. de Valença, Iñigo Virto, Adrian A. Wackett, Matthew W. Warren, Emily R. Webster, Nathaniel H. Wehr, Joann K. Whalen, Michael B. Wironen, Volkmar Wolters, Pengfei Wu, Irina V Zenkova, Weixin Zhang, Erin K. Cameron, Nico Eisenhauer, Helen R. P. Phillips, Elizabeth M. Bach, Marie L. C. Bartz, Joanne M. Bennett, Rémy Beugnon, Maria J. I Briones, George G. Brown, Olga Ferlian, Konstantin B. Gongalsky, Carlos A. Guerra, Birgitta König-Ries, Julia J. Krebs, Alberto Orgiazzi, Kelly S. Ramirez, David J. Russell, Benjamin Schwarz, Diana H. Wall, Ulrich Brose, Thibaud Decaëns, Patrick Lavelle, Michel Loreau, Jérôme Mathieu, Christian Mulder, Wim H. van der Putten, Matthias C. Rillig, Madhav P. Thakur, Franciska T. de Vries, David A. Wardle, Christian Ammer, Sabine Ammer, Miwa Arai, Fredrick O. Ayuke, Geoff H. Baker, Dilmar Baretta, Dietmar Barkusky, Robin Beauséjour, Jose C. Bedano, Klaus Birkhofer, Eric Blanchart, Bernd Blossey, Thomas Bolger, Robert L. Bradley, Michel Brossard, James C. Burtis, Yvan Capowiez, Timothy R. Cavagnaro, Amy Choi, Julia Clause, Daniel Cluzeau, Anja Coors, Felicity V Crotty, Jasmine M. Crumsey, Andrea Dávalos, Darío J. Díaz Cosín, Annise M. Dobson, Anahí Domínguez, Andrés Esteban Duhour, Nick van Eekeren, Christoph Emmerling, Liliana B. Falco, Rosa Fernández, Steven J. Fonte, Carlos Fragoso, André L. C. Franco, Abegail Fusilero, Anna P. Geraskina, Shaieste Gholami, Grizelle Gonzalez, Michael J. Gundale, Mónica Gutiérrez López, Branimir K. Hackenberger, Davorka K. Hackenberger, Luis M. Hernández, Jeff R. Hirth, Takuo Hishi, Andrew R. Holdsworth, Martin Holmstrup, Kristine N. Hopfensperger, Esperanza Huerta Lwanga, Veikko Huhta, Tunsisa T. Hurisso, Basil V Iannone, Madalina Iordache, Ulrich Irmler, Mari Ivask, Juan B. Jesús, Jodi L. Johnson-Maynard, Monika Joschko, Nobuhiro Kaneko, Radoslava Kanianska, Aidan M. Keith, Maria L. Kernecker, Armand W. Koné, Yahya Kooch, Sanna T. Kukkonen, H. Lalthanzara, Daniel R. Lammel, Iurii M. Lebedev, Edith Le Cadre, Noa K. Lincoln, Danilo López-Hernández, Scott R. Loss, Raphael Marichal, Radim Matula, Yukio Minamiya, Jan Hendrik Moos, Gerardo Moreno, Alejandro Morón-Ríos, Hasegawa Motohiro, Bart Muys, Johan Neirynck, Lindsey Norgrove, Marta Novo, Visa Nuutinen, Victoria Nuzzo, P. Mujeeb Rahman, Johan Pansu, Shishir Paudel, Guénola Pérès, Lorenzo Pérez-Camacho, Jean-François Ponge, Jörg Prietzel, Irina B. Rapoport, Muhammad Imtiaz Rashid, Salvador Rebollo, Miguel Á. Rodríguez, Alexander M. Roth, Guillaume X. Rousseau, Anna Rozen, Ehsan Sayad, Loes van Schaik, Bryant Scharenbroch, Michael Schirrmann, Olaf Schmidt, Boris Schröder, Julia Seeber, Maxim P. Shashkov, Jaswinder Singh. 2021. Global data on earthworm abundance, biomass, diversity and corresponding environmental properties
- Jess K. Zimmerman, Tana E. Wood, Grizelle Gonzalez, Alonso Ramirez, Whendee L. Silver, Maria Uriarte, Michael R. Willig, Robert B. Waide, Ariel Lugo. 2021. Disturbance and resilience in the Luquillo Experimental Forest
- N.M. Soto-Parrilla, Tamara Heartsill Scalley. 2021. Missing the Trees for the Forest: Post-Hurricane Understory Vegetation in Relation to Spatial Variation
- M.C. Leon, Tamara Heartsill Scalley, I. Santiago, W. H. McDowell. 2021. Hydrological Mapping in the Luquillo Experimental Forest: New Local Datum Improves Watershed Ecological Knowledge
- Smruthi Karthikeyan, Luis H. Orellana, Eric R. Johnston, Janet K. Hatt, Frank E. Löffler, Héctor L. Ayala-del-Río, Grizelle Gonzalez, Konstantinos T. Konstantinidis. 2021. Metagenomic Characterization of Soil Microbial Communities in the Luquillo Experimental Forest (Puerto Rico) and Implications for Nitrogen Cycling
- Rosanise Odell, Virginia Rose Seagal, Caroline May, Tanya Hawley Matlaga, Patricia Burrowes, Tana E. Wood. 2020. Coqui diet note
- Robert Muscarella, Samira Kolyaie, Douglas C. Morton, Jess K. Zimmerman, María Uriarte. 2020. Effects of topography on tropical forest structure depend on climate context
- Alberto Pastor Ibáñez. 2020. Caracterización post-huracán de la vegetación ribereña en los arroyos de cabecera de un bosque tropical montano-bajo; estructura arbórea y composición de especies
- Kelsey R. Carter, Tana E. Wood, Sasha C. Reed, Elsa C. Schwartz, Madeline B. Reinsel, Xi Yang, Molly A. Cavaleri. 2020. Photosynthetic and Respiratory Acclimation of Understory Shrubs in Response to in situ Experimental Warming of a Wet Tropical Forest
- Elvis Torres Delgado. 2020. Cloud and aerosol properties under the Influence of different air masses
- Wesley X. Torres-Pérez. 2020. Toxicity of Organophosphate and Pyrethroids Pesticides On The Tropical Freshwater Shrimp Xiphocaris Elongata
- Claudia J. Garnica-Díaz. 2020. Plant functional diversity across two elevational gradients in serpentine and volcanic soils of Puerto Rico
- Deborah K. Kennard, David Matlaga, Joanne Sharpe, Clay King, Aura M. Alonso‐Rodríguez, Sasha C. Reed, Molly A. Cavaleri, Tana E. Wood. 2020. Tropical understory herbaceous community responds more strongly to hurricane disturbance than to experimental warming
- Claudia M. Ospina-Sánchez, José G. Palacios-Vargas, Grizelle Gonzalez. 2020. A new Caribbean species of Hylaeanura Arlé, 1966 (Collembola, Neanuridae, Pseudachorutinae)
- Claudia Ospina-Sánchez, Felipe Soto-Adames, Grizelle Gonzalez. 2020. Checklist and distribution of Collembola from Greater Puerto Rico
- Iana F. Grullón – Penkova, Jess K. Zimmerman, Grizelle Gonzalez. 2020. Green roofs in the tropics: design considerations and vegetation dynamics
- Ashley Van Beusekom, Grizelle Gonzalez, Sarah Stankavich, Jess K. Zimmerman, Alonso Ramírez. 2020. Understanding tropical forest abiotic response to hurricanes using experimental manipulations, field observations, and satellite data
- Ariel Lugo, Oscar J. A. Martínez, Ernesto Medina, Gerardo Aymard, Tamara Heartsill Scalley. 2020. Novelty in the tropical forests of the 21st century
- Sasha C. Reed, Robin Reibold, Molly A Cavaleri, Aura. M. Alonso-Rodríguez, Megan Berberich, Tana E. Wood. 2020. Chapter Six - Soil biogeochemical responses of a tropical forest to warming and hurricane disturbance
- Daniela Yaffar, Richard J. Norby. 2020. A historical and comparative review of 50 years of root data collection in Puerto Rico
- Benedicte Bachelot, Aura M. Alonso-Rodríguez, Laura Aldrich-Wolfe, Molly A. Cavaleri, Sasha C. Reed, Tana E. Wood. 2020. Altered climate leads to positive density-dependent feedbacks in a tropical wet forest
- Claudia M. Ospina-Sánchez, José G. Palacios-Vargas, Grizelle Gonzalez. 2020. New species of Furculanurida (Collembola, Neanuridae, Pseudachorutinae) from the Luquillo Mountains, Puerto Rico
- James B. Shanley, Mark Marvin-DiPasquale, Oksana Lane, Wayne J. Arendt, Steven Hall, William H. McDowell. 2019. Resolving a paradox—high mercury deposition, but low bioaccumulation in northeastern Puerto Rico
- Ivia I. Moreno Rosado. 2019. Diversity and dynamics of arthropods and microbes, and nutrient release during green litter decomposition in a simulated hurricane experiment
- Claudia M. Ospina-Sánchez. 2019. Role of Microhabitats and environment variation on Collembola (Hexapoda: Entognatha) populations in the Luquillo Experimental Forest: a montane environment
- Kelsey R Carter. 2019. Ecophysiological responses of tropical woody species to ambient and elevated temperatures
- Tamara Heartsill Scalley, Gerónimo Mercado. 2019. Sound Circles/Inmersion
- Grizelle Gonzalez. 2019. The Brain and the Body
- M.A. McClintock, W.H. McDowell, Grizelle Gonzalez, Michael Schulz, J.C. Pett-Ridge. 2019. African dust deposition in Puerto Rico: Analysis of a 20-year rainfall chemistry record and comparison with models
- Jessica Chappell, S. Kyle McKay, Mary C. Freeman, Catherine M. Pringle. 2019. Long‐term (37 years) impacts of low‐head dams on freshwater shrimp habitat connectivity in northeastern Puerto Rico
- Azad Henareh Khalyani, William A. Gould, Michael J. Falkowski, Robert Muscarella, María Uriarte, Foad Yousef. 2019. Climate change increases potential plant species richness on Puerto Rican uplands
- Tania López-Marrero, Tamara Heartsill Scalley, Carlos F. Rivera-López, Isabel A. Escalera-García, Mariangelí Echevarría-Ramos. 2019. Broadening Our Understanding of Hurricanes and Forests on the Caribbean Island of Puerto Rico: Where and What Should We Study Now?
- Ariel Lugo. 2019. Acta Científica 31(1-3)
- M. R. Willig, L. Woolbright, S. J. Presley, T. D. Schowalter, R. B. Waide, Tamara Heartsill Scalley, J. K. Zimmerman, Grizelle Gonzalez, Ariel Lugo. 2019. Populations are not declining and food webs are not collapsing at the Luquillo Experimental Forest
- Tana E. Wood, Grizelle Gonzalez, Whendee Silver, Sasha Reed, Molly Cavaleri. 2019. On the Shoulders of Giants: Continuing the Legacy of Large-Scale Ecosystem Manipulation Experiments in Puerto Rico
- Ariel Lugo, Grizelle Gonzalez. 2019. Introduction to the Special Issue on Tropical Forests: Management and Ecology in the Anthropocene
- Omar Gutiérrez del Arroyo, Whendee L. Silver. 2018. Disentangling the long‐term effects of disturbance on soil biogeochemistry in a wet tropical forest ecosystem
- Kaitlin J. Farrell, Amy D. Rosemond, John S. Kominoski, Sophia M. Bonjour, Janine Rüegg, Lauren E. Koenig, Christina L. Baker, Matt T. Trentman, Tamara K. Harms, William H. McDowell. 2018. Variation in Detrital Resource Stoichiometry Signals Differential Carbon to Nutrient Limitation for Stream Consumers Across Biomes
- Ariel Lugo. 2018. Characterization of ten extreme disturbance events in the context of social and ecological systems
- Paul W. Miller, Thomas L. Mote, Craig A. Ramseyer, Ashley Van Beusekom, Martha Scholl, Grizelle Gonzalez. 2018. A 42 year inference of cloud base height trends in the Luquillo Mountains of northeastern Puerto Rico
- Ashley Van Beusekom, Nora Alvarez-Berrios, William A. Gould, Maya Quinones, Grizelle Gonzalez. 2018. Hurricane Maria in the U.S. Caribbean: Disturbance Forces, Variation of Effects, and Implications for Future Storms
- Xiaolin Zhu, Eileen Helmer. 2018. An automatic method for screening clouds and cloud shadows in optical satellite image time series in cloudy regions
- Ariel Lugo, Carlos Rodriguez, Ian Fermont, Ivan Vicens. 2018. Response to Hurricanes of Pinus caribaea var hondurensis Plantations in Puerto Rico
- Alida Mau, Sasha Reed, Tana E. Wood, Molly Cavaleri. 2018. Temperate and Tropical Forest Canopies are Already Functioning beyond Their Thermal Thresholds for Photosynthesis
- Benedicte Bachelot, María Uriarte, Robert Muscarella, Jimena Forero-Montaña, Jill Thompson, Krista McGuire, Jess Zimmerman, Nathan G. Swenson, James S. Clark. 2018. Associations among arbuscular mycorrhizal fungi and seedlings are predicted to change with tree successional status
- Xianbin Liu, Xiucheng Zeng, Xiaoming Zou, D. Jean Lodge, Sarah Stankavich, Grizelle Gonzalez, Sharon Cantrell. 2018. Responses of Soil Labile Organic Carbon to a Simulated Hurricane Disturbance in a Tropical Wet Forest
- Xianbin Liu, Xiucheng Zeng, Xiaoming Zou, Grizelle Gonzalez, Chao Wang, Si Yang. 2018. Litterfall Production Prior to and during Hurricanes Irma and Maria in Four Puerto Rican Forests
- Barbara Richardson, Michael Richardson, Grizelle Gonzalez. 2018. Responses of Two Litter-Based Invertebrate Communities to Changes in Canopy Cover in a Forest Subject to Hurricanes
- Maya Quinones, Isabel K. Parés-Ramos, William A. Gould, Grizelle Gonzalez, Kathleen McGinley, Pedro Ríos. 2018. El Atlas del Bosque Nacional El Yunque
- Maya Quinones, Isabel K. Parés-Ramos, William A. Gould, Grizelle Gonzalez, Kathleen McGinley, Pedro Ríos. 2018. El Yunque National Forest Atlas
- Chelse M. Prather, Gary E. Belovsky, Sharon A. Cantrell, Grizelle Gonzalez. 2018. Tropical herbivorous phasmids, but not litter snails, alter decomposition rates by modifying litter bacteria
- André-Jean Francez, Andrea Fischer, Andreas Bohner, Andrey Malyshev, Andrijana Andrić, Andy Smith, Angela Stanisci, Anikó Seres, Anja Schmidt, Anna Avila, Anne Probst, Annie Ouin, Anzar A. Khuroo, Arne Verstraeten, Arely N. Palabral-Aguilera, Artur Stefanski, Aurora Gaxiola, Bart Muys, Bernard Bosman, Bernd Ahrends, Bill Parker, Birgit Sattler, Bo Yang, Bohdan Juráni, Brigitta Erschbamer, Carmen Eugenia Rodriguez Ortiz, Casper T. Christiansen, E. Carol Adair, Céline Meredieu, Cendrine Mony, Charles A. Nock, Chi-Ling Chen, Chiao-Ping Wang, Christel Baum, Christian Rixen, Christine Delire, Christophe Piscart, Christopher Andrews, Corinna Rebmann, Cristina Branquinho, Dana Polyanskaya, David Fuentes Delgado, Dirk Wundram, Diyaa Radeideh, Eduardo Ordóñez-Regil, Edward Crawford, Elena Preda, Elena Tropina, Elli Groner, Eric Lucot, Erzsébet Hornung, Esperança Gacia, Esther Lévesque, Evanilde Benedito, Evgeny A. Davydov, Evy Ampoorter, Fabio Padilha Bolzan, Felipe Varela, Ferdinand Kristöfel, Fernando T. Maestre, Florence Maunoury-Danger, Florian Hofhansl, Florian Kitz, Flurin Sutter, Francisco Cuesta, Francisco de Almeida Lobo, Franco Leandro de Souza, Frank Berninger, Franz Zehetner, Georg Wohlfahrt, George Vourlitis, Geovana Carreño-Rocabado, Gina Arena, Gisele Daiane Pinha, Grizelle Gonzalez, Authors Additional, Ian Yesilonis, Authors Additional, Lindsey Rustad, al. et, Ika Djukic, Sebastian Kepfer-Rojas, Inger Kappel Schmidt, Ana Carolina Ruiz Fernández, Ana I. Sousa, Ana I. Lillebø, Klaus Steenberg Larsen, Claus Beier, Björn Berg, Kris Verheyen, Adriano Caliman, Alain Paquette, Alba Gutiérrez-Girón, Alberto Humber, Alejandro Valdecantos, Alessandro Petraglia, Heather Alexander, Algirdas Augustaitis, Amélie Saillard, Anderson da Rocha Gripp. 2018. Early stage litter decomposition across biomes
- Xiaohui Feng, María Uriarte, Grizelle Gonzalez, Sasha Reed, Jill Thompson, Jess K. Zimmerman, Lora Murphy. 2018. Improving predictions of tropical forest response to climate change through integration of field studies and ecosystem modeling
- Aura M. Alonso-Rodríguez, Molly A. Cavaleri, Sasha C. Reed, Grizelle Gonzalez, Tana E. Wood, Bruce A. Kimball. 2018. Infrared heater system for warming tropical forest understory plants and soils
- Kathleen McGinley, Guy Robertson, Kathleen Friday, Constance Carpenter. 2017. Assessing Forest Sustainability in the Tropical Islands of the United States
- Marconi Campos-Cerqueira, Wayne J. Arendt, Joseph M. Wunderle Jr., T. Mitchell. Aide. 2017. Have bird distributions shifted along an elevational gradient on a tropical mountain?
- Grizelle Gonzalez, María F. Barberena. 2017. Ecology of soil arthropod fauna in tropical forests: A review of studies from Puerto Rico
- Ashley Van Beusekom, Grizelle Gonzalez, Maria Rivera. 2017. Reply to “Comments on ‘Short-Term Precipitation and Temperature Trends along an Elevation Gradient in Northeastern Puerto Rico’ ”
- Sheila F. Murphy, Robert F. Stallard, Martha A. Scholl, Grizelle Gonzalez, Angel J. Torres-Sánchez. 2017. Reassessing rainfall in the Luquillo Mountains, Puerto Rico: Local and global ecohydrological implications
- Ashley Van Beusekom, Grizelle Gonzalez, Martha A. Scholl. 2017. Analyzing cloud base at local and regional scales to understand tropical montane cloud forest vulnerability to climate change
- Tamara Heartsill Scalley. 2017. Insights on Forest Structure and Composition from Long-Term Research in the Luquillo Mountains
- Dingfang Chen, Mei Yu, Grizelle Gonzalez, Xiaoming Zou, Qiong Gao. 2017. Climate impacts on soil carbon processes along an elevation gradient in the tropical Luquillo Experimental Forest
- James Aaron Hogan, Jess K. Zimmerman, Jill Thompson, Christopher J. Nytch, María Uriarte. 2016. The interaction of land-use legacies and hurricane disturbance in subtropical wet forest: twenty-one years of change
- Ching-Yu Huang, Grizelle Gonzalez, Paul F. Hendrix. 2016. Resource Utilization by Native and Invasive Earthworms and Their Effects on Soil Carbon and Nitrogen Dynamics in Puerto Rican Soils
- D. Jean Lodge, Dirk Winter, Grizelle Gonzalez, Naomi Clum. 2016. Effects of hurricane-felled tree trunks on soil carbon, nitrogen, microbial biomass, and root length in a wet tropical forest
- Ariel Lugo, J.L. Frangi. 2016. Long-term response of Caribbean palm forests to hurricanes
- James W. Dalling, Katherine Heineman, Grizelle Gonzalez, Rebecca Ostertag. 2016. Geographic, environmental and biotic sources of variation in the nutrient relations of tropical montane forests
- Marcela Zalamea, Grizelle Gonzalez, D. Jean Lodge. 2016. Physical, Chemical, and Biological Properties of Soil under Decaying Wood in a Tropical Wet Forest in Puerto Rico
- Nicholas J. Bouskill, Tana E. Wood, Richard Baran, Zaw Ye, Benjamin P. Bowen, HsiaoChien Lim, Jizhong Zhou, Joy D. Van Nostrand, Peter Nico, Trent R. Northen, Whendee L. Silver, Eoin L. Brodie. 2016. Belowground Response to Drought in a Tropical Forest Soil. I. Changes in Microbial Functional Potential and Metabolism
- Nicholas J. Bouskill, Tana E. Wood, Richard Baran, Zhao Hao, Zaw Ye, Ben P. Bowen, Hsiao Chien Lim, Peter S. Nico, Hoi-Ying Holman, Benjamin Gilbert, Whendee L. Silver, Trent R. Northen, Eoin L. Brodie. 2016. Belowground Response to Drought in a Tropical Forest Soil. II. Change in Microbial Function Impacts Carbon Composition
- Tana E. Wood, Danielle Matthews, Karen Vandecar, Deborah Lawrence. 2016. Short-term variability in labile soil phosphorus is positively related to soil moisture in a humid tropical forest in Puerto Rico
- Ariel Lugo. 2015. Acta Científica Vol. 29 (1-3)
- Martha A. Scholl, James B. Shanley, Sheila F. Murphy, Jane K. Willenbring, Grizelle Gonzalez. 2015. Stable-isotope and solute-chemistry approaches to flow characterization in a forested tropical watershed, Luquillo Mountains, Puerto Rico
- Aaron Shiels, Grizelle Gonzalez. 2015. Tropical Forest Responses to Large-Scale Experiments
- Ashley Van Beusekom, Grizelle Gonzalez, Maria Rivera. 2015. Short-term precipitation and temperature trends along an elevation gradient in northeastern Puerto Rico
- Gavin McNicol, Whendee L. Silver. 2014. Separate effects of flooding and anaerobiosis on soil greenhouse gas emissions and redox sensitive biogeochemistry
- Tania del M. Lopez-Marrero, Tamara Heartsill Scalley. 2014. Land-cover composition, water resources and land management in the watersheds of the Luquillo Mountains, northeastern Puerto Rico.
- Michael R. Willig, Christopher P. Bloch, Steven J. Presley. 2014. Experimental decoupling of canopy opening and debris addition on tropical gastropod populations and communities
- Jess K. Zimmerman, James Aaron Hogan, Aaron B. Shiels, John E. Bithorn, Samuel Matta Carmona, Nicholas Brokaw. 2014. Seven-year responses of trees to experimental hurricane effects in a tropical rainforest, Puerto Rico
- Joanne M. Sharpe, Aaron B. Shiels. 2014. Understory fern community structure, growth and spore production responses to a large-scale hurricane experiment in a Puerto Rico rainforest
- Timothy D. Schowalter, Michael R. Willig, Steven J. Presley. 2014. Canopy arthropod responses to experimental canopy opening and debris deposition in a tropical rainforest subject to hurricanes
- Chelse Prather. 2014. Divergent responses of leaf herbivory to simulated hurricane effects in a rainforest understory
- William H. McDowell, Daniel Liptzin. 2014. Linking soils and streams: Response of soil solution chemistry to simulated hurricane disturbance mirrors stream chemistry following a severe hurricane
- Paul D. Klawanski, Ben Dalton, Aaron B. Shiels. 2014. Coqui frog populations are negatively affected by canopy opening but not detritus deposition following an experimental hurricane in a tropical rainforest
- W.L. Silver, S.J. Hall, Grizelle Gonzalez. 2014. Differential effects of canopy trimming and litter deposition on litterfall and nutrient dynamics in a wet subtropical forest
- Grizelle Gonzalez, M.R. Willig, A.B. Shiels. 2014. Responses to canopy loss and debris deposition in a tropical forest ecosystem: Synthesis from an experimental manipulation simulating effects of hurricane disturbance
- A.B. Shiels, Grizelle Gonzalez. 2014. Understanding the key mechanisms of tropical forest responses to canopy loss and biomass deposition from experimental hurricane effects
- Sharon A. Cantrell, Marirosa Molina, D. Jean Lodge, Francisco J. Rivera-Figueroa, Maria Ortiz, Albany A. Marchetti, Mike J. Cyterski, José R. Pérez-Jiménez. 2014. Effects of a simulated hurricane disturbance on forest floor microbial communities
- D. Jean Lodge, Sharon A. Cantrell, Grizelle Gonzalez. 2014. Effects of canopy opening and debris deposition on fungal connectivity, phosphorus movement between litter cohorts and mass loss
- Ariel Lugo, Tamara Heartsill Scalley. 2014. Research in the Luquillo Experimental Forest has advanced understanding of tropical forests and resolved management issues
- W.H. McDowell, R.L. Brereton, F.N. Scatena, J.B. Shanley, N.V. Brokaw, Ariel Lugo. 2013. Interactions between lithology and biology drive the long-term response of stream chemistry to major hurricanes in a tropical landscape
- Sharon A. Cantrell, D. Jean Lodge, Carlos A. Cruz, Luis M. García, Jose R. Pérez-Jiménez, Marirosa Molina. 2013. Differential abundance of microbial functional groups along the elevation gradient from the coast to the Luquillo Mountains
- R. B. Waide, D. E. Comarazamy, J. E. González, C. A. S. Hall, Ariel Lugo, J. C. Luvall, D. J. Murphy, J. R. Ortiz-Zayas, N. D. Ramírez-Beltrán, F. N. Scatena, W. L. Silver. 2013. Climate variability at multiple spatial and temporal scales in the Luquillo Mountains, Puerto Rico
- P. L. Weaver, William A. Gould. 2013. Forest vegetation along environmental gradients in northeastern Puerto Rico
- M. R. Willig, S. J. Presley, C. P. Bloch, J. Alvarez, NO-VALUE. 2013. Population, community, and metacommunity dynamics of terrestrial gastropods in the Luquillo Mountains: a gradient perspective
- E. O. Vázquez Plass, Joseph M. Wunderle Jr. 2013. Avian distribution along a gradient of urbanization in northeastern Puerto Rico
- A. B. Shiels, L. R. Walker, NO-VALUE. 2013. Landslides cause spatial and temporal gradients at multiple scales in the Luquillo Mountains of Puerto Rico
- W. L. Silver, D. Liptzin, M. Almaraz, NO-VALUE. 2013. Soil redox dynamics and biogeochemistry along a tropical elevation gradient
- B. A. Richardson, M. J. Richardson, NO-VALUE. 2013. Litter-based invertebrate communities in forest floor and bromeliad microcosms along an elevational gradient in Puerto Rico
- C.-L. Ping, G. J. Michaelson, C. A. Stiles, Grizelle Gonzalez. 2013. Soil characteristics, carbon stores, and nutrient distribution in eight forest types along an elevational gradient, eastern Puerto Rico
- E. Medina, Grizelle Gonzalez, Maria Rivera. 2013. Spatial and temporal heterogeneity of rainfall inorganic ion composition in northeastern Puerto Rico
- Ariel Lugo. 2013. Foreword, up, down, and across the mountains: a new look at the Luquillo Mountains
- W. G. Lash-Marshall, C. A. S. Hall, W. W. Fu, NO-VALUE. 2013. Using gradient analysis to simulate the spatial structure and function of the Luquillo Experimental Forest
- N. L. Harris, E. Medina. 2013. Changes in leaf properties across an elevation gradient in the Luquillo Mountains, Puerto Rico
- N. L. Harris, C. A. S. Hall, Ariel Lugo. 2013. A test of the maximum power hypothesis along an elevational gradient in the Luquillo Mountains of Puerto Rico
- A. Gioda, O.L. Mayol-Bracero, F. N. Scatena, K. C. Weathers, V. L. Mateus, W. H. McDowell. 2013. Chemical constituents in clouds and rainwater in the Puerto Rican rainforest: potential sources and seasonal drivers
- A. C. Gellis, NO-VALUE. 2013. Factors influencing storm-generated suspended-sediment concentrations and loads in four basins of contrasting land use, humid-tropical Puerto Rico
- Grizelle Gonzalez, M.R. Willig, R.B. Waide. 2013. Ecological gradient analyses in a tropical landscape: multiple perspectives and emerging themes
- Grizelle Gonzalez, R.B. Waide, M.R. Willig. 2013. Advancements in the understanding of spatiotemporal gradients in tropical landscapes: a Luquillo focus and global perspective
- Grizelle Gonzalez, M.M. Luce. 2013. Woody debris characterization along an elevation gradient in northeastern Puerto Rico
- D. F. Cusack. 2013. Soil nitrogen levels are linked to decomposition enzyme activities along an urban-remote tropical forest gradient
- Nicholas J. Bouskill, Hsiao Chien Lim, Sharon Borglin, Rohit Salve, Tana E. Wood, Whendee L. Silver, Eoin L. Brodie. 2013. Pre-exposure to drought increases the resistance of tropical forest soil bacterial communities to extended drought
- Wayne J. Arendt, S.S. Qian, K. Mineard. 2013. Population decline of the Elfin-woods Warbler Setophaga angelae in eastern Puerto Rico
- A. Argerich, Sherri Johnson, Stephen D. Sebestyen, Charles C. Rhoades, E. Greathouse, Jennifer D. Knoepp, Mary Beth Adams, G.E. Likens, John Campbell, W.H. McDowell, F.N. Scatena, G.G. Ice. 2013. Trends in stream nitrogen concentrations for forested reference catchments across the USA
- Peter L. Weaver. 2012. La sierra de Luquillo: Los recursos forestales y su historia
- Tamara Heartsill Scalley. 2012. Vegetation
- Tana E. Wood, W. L. Silver. 2012. Strong spatial variability in trace gas dynamics following experimental drought in a humid tropical forest
- P. L. Weaver. 2012. The Luquillo Mountains: forest resources and their history
- M.R. Willig, C.P. Bloch, A.P. Covich, C.A.S. Hall, D. Jean Lodge, Ariel Lugo, W.L. Silver, R.B. Waide, L.R. Walker, J.K. Zimmerman. 2012. Long-term research in the Luquillo Mountains, synthesis and foundations for the future
- F.N. Scatena, J.F. Blanco, K.H. Beard, R.B. Waide, Ariel Lugo, N. Brokaw, W.L. Silver, B.L. Haines, J.K. Zimmerman. 2012. Disturbance regime
- W.H. McDowell, F.N. Scatena, R.B. Waide, N. Brokaw, G.R. Camilo, A.P. Covich, T.A. Crowl, Grizelle Gonzalez, E.A. Greathouse, P. Klawinski, D. Jean Lodge, Ariel Lugo, C.M. Pringle, B.A. Richardson, M.J. Richardson, D.A. Schaefer, W.L. Silver, J. Thompson, D.J. Vogt, K.A. Vogt, M.R. Willig, L.L. Woolbright, X. Zou, J.K. Zimmerman. 2012. Geographic and ecological setting of the Luquillo Mountains
- Ariel Lugo, F.N. Scatena, R.B. Waide, E.A. Greathouse, C.M. Pringle, M.R. Willig, K.A. Vogt, L.R. Walker, Grizelle Gonzalez, W.H. McDowell, J. Thompson, NO-VALUE. 2012. Management implications and applications of long-term ecological research
- Ariel Lugo, R.B. Waide, M.R. Willig, T.A. Crowl, F.N. Scatena, J. Thompson, W.L. Silver, W.H. McDowell, N. Brokaw. 2012. Ecological paradigms for the tropics: old questions and continuing challenges
- F. Holwerda, L.A. Bruijnzeel, F.N. Scatena, H.F. Vugts, A.G.C.A. Meesters. 2012. Wet canopy evaporation from a Puerto Rican lower montane rain forest: the importance of realistically estimated aerodynamic conductance
- S. Martinuzzi, I.K. Páres-Ramos, William A. Gould. 2012. Land use, population dynamics, and land-cover change in Eastern Puerto Rico
- Marcela Zalamea, Grizelle Gonzalez, William A. Gould. 2012. Comparing litterfall and standing vegetation: assessing the footprint of litterfall traps
- Nancy L. Harris, Ariel Lugo, Sandra Brown, Tamara Heartsill Scalley. 2012. Luquillo Experimental Forest: Research History and Opportunities
- D.A. Wall, Grizelle Gonzalez, B.L. Simmons. 2011. Seasonally dry forest soil biodiversity and functioning
- D. J. Murphy, M. H. Hall, C. A. S. Hall, G. M. Heisler, S. V. Stehman, C. Anselmi-Molina, NO-VALUE. 2011. The relationship between land cover and the urban heat island in northeastern Puerto Rico
- L.R. Reitsma, M.T. Hallworth, K. Parent. 2011. Habitat use of the Louisiana Waterthrush during the non-breeding season in Puerto Rico
- Alejandro A. Royo, Tamara Heartsill Scalley, Samuel Moya, Fred N. Scatena. 2011. Non-arborescent vegetation trajectories following repeated hurricane disturbance: ephemeral versus enduring responses
- Joseph M. Wunderle Jr., Wayne J. Arendt, Jose William Beltran. 2010. Changes in home range of breeding and post-breeding male Pearly-eyed Thrashers in the Luquillo Mountains of Puerto Rico
- Tamara Heartsill Scalley, Frederick N. Scatena, Ariel Lugo, Samuel Moya, Carlos R. Estrada Ruiz. 2010. Changes in structure, composition, and nutrients during 15 years of hurricane-induced succession in a subtropical wet forest in Puerto Rico
- Frank Wadsworth. 2009. Topographic frequency of trees in the tabonuco forest of the Luquillo Mountains
- Frank Wadsworth. 2009. Early assessment of the trees of the Luquillo Mountains
- Tamara Heartsill Scalley, Todd A. Crowl, Jill Thompson. 2009. Tree species distributions in relation to stream distance in mid-montane wet forest, Puerto Rico
- Grizelle Gonzalez, Emerita Garcia, Veronica Cruz, Sonia Borges, Marcela Zalamea, Maria Rivera. 2007. Earthworm communities along an elevation gradient in Northeastern Puerto Rico.
- Rebecca Ostertag, Whendee L. Silver, Ariel Lugo. 2005. Factors Affecting Mortality and Resistance to Damage Following Hurricanes in a Rehabilitated Subtropical Moist Forest
- Wayne J. Arendt. 2004. Open- and cavity-nesting traits merge in the evolution of an avian supertramp, the Pearly-eyed Thrasher (Margarops fuscatus).
- Mark I Cook, Steven R. Beissinger, Gary A. Toranzos, Roberto A. Rodriguez, Wayne J. Arendt. 2003. Trans–shell infection by pathogenic micro–organisms reduces the shelf life of non–incubated bird's eggs: a constraint on the onset of incubation?
- P.L. Weaver. 2002. A chronology of hurricane induced changes in Puerto Rico’s Lower Montane Rain Forest
- P.L. Weaver. 1999. Impacts of Hurricane Hugo on the Dwarf Cloud Forest of Puerto Rico’s Luquillo Mountains
- Jorge L. Frangi, Ariel Lugo. 1998. A flood plain palm forest in the Luquillo Mountains of Puerto Rico five years after Hurricane Hugo
- L.R. Walker, J.K. Zimmerman, D.J. Lodge, S. Guzmán-Grajales. 1996. An altitudinal comparison of growth and species composition in hurricane-damaged forests in Puerto Rico
- Joseph M. Wunderle Jr. 1995. Responses of bird populations in a Puerto Rican forest to Hurricane Hugo: The first 18 months
- Marcia H. Wilson, Cameron B. Kepler, Noel F.R. Snyder, Scott R. Derrickson, F. Josh Dein, James W. Wiley, Joseph M. Wunderle Jr., Ariel Lugo, David L. Graham, William D. Toone. 1994. Puerto Rican Parrots and Potential Limitations of the Metapopulation Approach to Species Conservation
- F.N. Scatena, W. Silver, T. Siccama, A. Johnson, M.J. Sanchez. 1993. Biomass and Nutrient Content of the Bisley Experimental Watersheds, Luquillo Experimental Forest, Puerto Rico, before and after Hurricane Hugo, 1989
- Khadga Basnet, Frederick N. Scatena, Gene E. Likens, Ariel Lugo. 1993. Ecological Consequences of Root Grafting in Tabonuco (Dacryodes excelsa) Trees in the Luquillo Experimental Forest, Puerto Rico
- Joseph M. Wunderle Jr. 1992. Sexual habitat segregation in Black-throated Blue Warblers wintering in Puerto Rico.
- Ariel Lugo, F.N. Scatena. 1992. Eipihytes and Climate Change Research in the Caribbean: A Proposal
- D. Jean Lodge, F. N. Scatena, C. E. Asbury, M. J. Sanchez. 1991. Fine Litterfall and Related Nutrient Inputs Resulting From Hurricane Hugo in Subtropical Wet and Lower Montane Rain Forests of Puerto Rico
- J.A. Parrotta, D.J. Lodge. 1991. Fine Root Dynamics in a Subtropical Wet Forest Following Hurricane Disturbance in Puerto Rico
- Jorge L. Frangi, Ariel E. Lugo. 1991. Hurricane Damage to a Flood Plain Forest in the Luquillo Mountains of Puerto Rico
- R.L. Sanford, W.J. Parton, D.S. Ojima, D.J. Lodge. 1991. Hurricane Effects on Soil Organic Matter Dynamics and Forest Production in the Luquillo Experimental Forest, Puerto Rico: Results of Simulation Modeling
- Karen H. Beard, Kristiina A. Vogt, Daniel J. Vogt, Frederick N. Scatena, Alan P. Covich, Ragnhildur Sigurdardottir, Thomas G. Siccama, Todd A. Crowl. 1991. Structural and functional responses of a subtropical forest to 10 years of hurricanes and droughts.
- Gerald D. Lindsey, Wayne J. Arendt, Jan Kalina, Grey Pendleton. 1991. Home range and movements of juvenile Puerto Rican Parrots
- Gerald D. Lindsey, Wayne J. Arendt. 1991. Radio Tracking Puerto Rican Parrots: Assessing triangulation accuracy in an insular rain forest
- P.L. Weaver. 1989. Forest changes after hurricanes in Puerto Rico’s Luquillo Mountains
- P.L. Weaver. 1986. Hurricane damage and recovery in the montane forests of the Luquillo Mountains of Puerto Rico
- Leslie S. Uhazy, Wayne J. Arendt. 1986. Pathogenesis associated with philornid myiasis (Diptera: Muscidae) on nestling Pearly-eyed Thrashers (Aves: Mimidae) in the Luquillo Rain Forest, Puerto Rico
- Wayne J. Arendt. 1985. Philornis ectoparasitism of Pearly-eyed Thrashers. I. Impact on growth and development of nestlings
- Wayne J. Arendt. 1985. Philornis ectoparasitism of Pearly-eyed Thrashers. II. Effects on adults and reproduction
- Thomas Crow. 1980. A Rainforest Chronicle: A 30-Year Record of Change in Structure and Composition at El Verde, Puerto Rico
- F.H. Wadsworth, G.H. Englerth. 1959. Effects of the 1956 hurricane on forests in Puerto Rico
- Jose Marrero. 1947. A survey of the forest plantations in the Caribbean National Forest