Stephen Matthews, PhD

Treesearch Publications

• Hannah L. Clipp, Kevin M. Potter, Matthew Peters, Anantha M. Prasad, Stephen Matthews, Andrew Gougherty. 2026. Warming temperatures and shifting precipitation patterns may exacerbate pest damage in North American forests
• Anantha M. Prasad, Matthew Peters, John Pedlar, Andrew Gougherty, Daniel McKenney, Franz Mora, Stephen Matthews, Steve McNulty, Lauro López‐Mata. 2025. Habitat and Migration Dynamics of North American Tree Species Under Climate Change
• Andrew Gougherty, Hannah L. Clipp, Anantha M. Prasad, Matthew Peters, Stephen Matthews. 2025. Integrating disturbance to improve our understanding of range-wide patterns in tree species abundance and demography
• Hannah L. Clipp, Christopher T. Rota, Stephen Matthews, Matthew Peters, Petra B. Wood. 2025. Potential resilience of forest birds in the Appalachian Mountains to future climate change during the breeding season
• Matthew Peters, Rebecca A. Rooney, Stephen Matthews, Anantha M. Prasad, Louis Iverson, Maria Janowiak. 2025. Climate Change Atlas user guide: version 1.0
• Andrew Gougherty, Anantha M. Prasad, Matthew Peters, Stephen Matthews, Bryce T. Adams. 2024. Climate Change and the Emergence of No‐Analog Forest Assemblages in North America
• Bryce T. Adams, Alejandro A. Royo, Christel C. Kern, Dustin Bronson, Stephen Matthews, Andrew Gougherty, Anantha M. Prasad, Louis Iverson, Evan Rehm, Patricia Raymond, Catherine Périé, Jessica R. Miesel, Amanda M. McGraw. 2024. Identifying climatically-compatible seedlots for the eastern US: building the predictive tools and knowledge to enable forest assisted migration
• Andrew Gougherty, Ashley Walters, Anantha M. Prasad, Matthew Peters, Stephen Matthews, Ian DeMerchant. 2024. Climate, Host Abundance and Spread: Unravelling the Drivers of Forest Pest Distributions in North America
• Aaron Skinner, Stephen Matthews, Michael Ward, Ian Souza-Cole, James Wright, Frank R. Thompson, Thomas Benson, Christopher M. Tonra. 2023. Eastern Whip-poor-wills have larger nonbreeding home ranges in areas with more agriculture and forest fragmentation
• Anantha M. Prasad, Matthew Peters, Stephen Matthews, Louis Iverson. 2023. Unpacking the 'black box': Improving ecological interpretation of regression‐based models
• James R. Wright, Stephen Matthews, Cornelia Wilson, Christopher M. Tonra. 2022. Preferences of avian seed-hoarders in advance of potential American chestnut reintroduction
• Alexander Fotis, Charles E. Flower, Jeff W. Atkins, Cornelia Wilson, Amanda D. Rodewald, Stephen Matthews. 2022. The short-term and long-term effects of honeysuckle removal on canopy structure and implications for urban forest management
• Matthew Peters, Stephen Matthews, Anantha M. Prasad, Louis Iverson. 2022. Defining landscape-level forest types: application of latent Dirichlet allocation to species distribution models
• Bryce T. Adams, Stephen Matthews. 2022. Feature-dependent group structures and hierarchical songbird-habitat relationships in a managed forest landscape
• Aaron A. Skinner, Michael P. Ward, Ian Souza‐Cole, James R. Wright, Frank R. Thompson, Thomas J. Benson, Stephen Matthews, Christopher M. Tonra. 2022. High spatiotemporal overlap in the non‐breeding season despite geographically dispersed breeding locations in the eastern whip‐poor‐will (Antrostomus vociferus)
• Bryce T. Adams, Stephen Matthews, Louis Iverson, Anantha M. Prasad, Matthew Peters, Kaiguang Zhao. 2021. Spring phenological variability promoted by topography and vegetation assembly processes in a temperate forest landscape
• Anantha M. Prasad, John Pedlar, Matthew Peters, Dan McKenney, Louis Iverson, Stephen Matthews, Bryce T. Adams. 2020. Combining US and Canadian forest inventories to assess habitat suitability and migration potential of 25 tree species under climate change
• Bryce T. Adams, Louis Iverson, Stephen Matthews, Matthew Peters, Anantha M. Prasad, David M. Hix. 2020. Mapping Forest Composition with Landsat Time Series: An Evaluation of Seasonal Composites and Harmonic Regression
• W. Keith Moser, Patricia Butler-Leopold, Constance Hausman, Louis Iverson, Todd A. Ontl, Leslie Brand, Stephen Matthews, Matthew Peters, Anantha M. Prasad. 2020. The impact of climate change on forest systems in the northern United States: Projections and implications for forest management [Chapter 8]
• Stephen Matthews, David M. Hix, James D. Palus, Erin E. Andrew, P. Charles Goebel, Donald Radcliffe. 2019. Assessing change in hardwood forests of the Wayne National Forest, southeastern Ohio
• Louis Iverson, Matthew Peters, Stephen Matthews, Anantha M. Prasad, Todd Hutchinson, Jarel Bartig, Joanne Rebbeck, Dan Yaussy, Susan L. Stout, Greg Nowacki. 2019. Adapting oak management in an age of ongoing mesophication but warming climate
• Louis Iverson, Anantha M. Prasad, Matthew Peters, Stephen Matthews. 2019. Facilitating Adaptive Forest Management under Climate Change: A Spatially Specific Synthesis of 125 Species for Habitat Changes and Assisted Migration over the Eastern United States
• Louis Iverson, Jarel L. Bartig, Gregory J. Nowacki, Matthew Peters, James M. Dyer, Todd Hutchinson, Stephen Matthews, Bryce T. Adams. 2019. USDA Forest Service section, subsection, and landtype descriptions for southeastern Ohio
• Matthew Peters, Louis Iverson, Anantha M. Prasad, Stephen Matthews. 2019. Utilizing the density of inventory samples to define a hybrid lattice for species distribution models: DISTRIB‐II for 135 eastern U.S. trees
• Louis Iverson, Matthew Peters, Anantha M. Prasad, Stephen Matthews. 2019. Analysis of Climate Change Impacts on Tree Species of the Eastern US: Results of DISTRIB-II Modeling
• James D. Palus, P. Charles Goebel, David M. Hix, Stephen Matthews. 2018. Structural and compositional shifts in forests undergoing mesophication in the Wayne National Forest, southeastern Ohio
• Patricia R. Butler-Leopold, Louis Iverson, Frank R. Thompson, Leslie Brandt, Stephen Handler, Maria Janowiak, P. Danielle Shannon, Chris Swanston, Scott Bearer, Alexander M. Bryan, Ken Clark, Greg Czarnecki, Philip DeSenze, William D. Dijak, Jacob S. Fraser, Paul F. Gugger, Andrea Hille, Justin Hynicka, Claire A. Jantz, Matthew C. Kelly, Katrina M. Krause, Inga La Puma, Deborah Landau, Richard G. Lathrop, Laura P. Leites, Evan Madlinger, Stephen Matthews, Gulnihal Ozbay, Matthew Peters, Anantha M. Prasad, David A. Schmit, Collin Shephard, Rebecca Shirer, Nicholas Skowronski, Al Steele, Susan L. Stout, Melissa Thomas-Van Gundy, John Thompson, Richard M. Turcotte, David A. Weinstein, Alfonso Yáñez. 2018. Mid-Atlantic forest ecosystem vulnerability assessment and synthesis: a report from the Mid-Atlantic Climate Change Response Framework project
• Louis Iverson, Matthew Peters, Jarel L. Bartig, Joanne Rebbeck, Todd Hutchinson, Stephen Matthews, Susan L. Stout. 2018. Spatial modeling and inventories for prioritizing investment into oak-hickory restoration
• Stephen Matthews, Louis Iverson, Matthew Peters, Anantha M. Prasad. 2018. Assessing potential climate change pressures across the conterminous United States: mapping plant hardiness zones, heat zones, growing degree days, and cumulative drought severity throughout this century
• Maria Janowiak, Anthony W. D'Amato, Chris Swanston, Louis Iverson, Frank R. Thompson, William D. Dijak, Stephen Matthews, Matthew Peters, Anantha M. Prasad, Jacob S. Fraser, Leslie Brandt, Patricia Butler-Leopold, Stephen Handler, P. Danielle Shannon, Diane Burbank, John Campbell, Charles Cogbill, Matthew Duveneck, Marla R. Emery, Nicholas Fisichelli, Jane Foster, Jennifer Hushaw, Laura S. Kenefic, Amanda Mahaffey, Toni Lyn Morelli, Nicholas J. Reo, Paul Schaberg, K. Rogers Simmons, Aaron Weiskittel, Sandy Wilmot, David Hollinger, Erin D. Lane, Lindsey Rustad, Pamela H. Templer. 2018. New England and northern New York forest ecosystem vulnerability assessment and synthesis: a report from the New England Climate Change Response Framework project
• Leslie Brandt, Abigail Derby Lewis, Lydia Scott, Lindsay Darling, Robert T. Fahey, Louis Iverson, David J. Nowak, Allison Bodine, Andrew Bell, Shannon Still, Patricia R. Butler, Andrea Dierich, Stephen Handler, Maria Janowiak, Stephen Matthews, Jason W. Miesbauer, Matthew Peters, Anantha M. Prasad, P. Danielle Shannon, Douglas Stotz, Chris Swanston. 2017. Chicago Wilderness region urban forest vulnerability assessment and synthesis: a report from the Urban Forestry Climate Change Response Framework Chicago Wilderness pilot project
• Stephen Matthews, Louis Iverson. 2017. Managing for delicious ecosystem service under climate change: can United States sugar maple (Acer saccharum) syrup production be maintained in a warming climate?
• Louis Iverson, Frank R. Thompson, Stephen Matthews, Matthew Peters, Anantha M. Prasad, William D. Dijak, Jacob S. Fraser, Wen J. Wang, Brice Hanberry, Hong He, Maria Janowiak, Patricia Butler, Leslie Brandt, Chris Swanston. 2017. Multi-model comparison on the effects of climate change on tree species in the eastern U.S.: results from an enhanced niche model and process-based ecosystem and landscape models
• Anantha M. Prasad, Louis Iverson, Stephen Matthews, Matthew Peters. 2016. A multistage decision support framework to guide tree species management under climate change via habitat suitability and colonization models, and a knowledge-based scoring system
• James S. Clark, Louis Iverson, Christopher W. W. Woodall, Craig D. Allen, David M. Bell, Don C. Bragg, Anthony W. D'Amato, Frank W. Davis, Michelle H. Hersh, Ines Ibanez, Stephen T. Jackson, Stephen Matthews, Neil Pederson, Matthew Peters, Mark W. Schwartz, Kristen M. Waring, Niklaus E. Zimmermann. 2016. The impacts of increasing drought on forest dynamics, structure, and biodiversity in the United States
• Louis Iverson, Kathleen Knight, Anantha M. Prasad, Daniel A. Herms, Stephen Matthews, Matthew Peters, Annemarie Smith, Diane M. Hartzler, Robert P. Long, John Almendinger. 2015. Potential species replacements for black ash (Fraxinus nigra) at the confluence of two threats: Emerald ash borer and a changing climate
• Patricia R. Butler, Louis Iverson, Frank R. Thompson, Leslie Brandt, Stephen Handler, Maria Janowiak, P. Danielle Shannon, Chris Swanston, Kent Karriker, Jarel Bartig, Stephanie Connolly, William D. Dijak, Scott Bearer, Steve Blatt, Andrea Brandon, Elizabeth Byers, Cheryl Coon, Tim Culbreth, Jad Daly, Wade Dorsey, David Ede, Chris Euler, Neil Gillies, David M. Hix, Catherine Johnson, Latasha Lyte, Stephen Matthews, Dawn McCarthy, Dave Minney, Daniel Murphy, Claire O’Dea, Rachel Orwan, Matthew Peters, Anantha M. Prasad, Cotton Randall, Jason Reed, Cynthia Sandeno, Thomas M. Schuler, Lesley Sneddon, Bill Stanley, Al Steele, Susan L. Stout, Randy Swaty, Jason Teets, Tim Tomon, Jim Vanderhorst, John Whatley, Nicholas Zegre. 2015. Central Appalachians forest ecosystem vulnerability assessment and synthesis: a report from the Central Appalachians Climate Change Response Framework project
• Matthew Peters, Louis Iverson, Stephen Matthews. 2015. Long-term droughtiness and drought tolerance of eastern US forests over five decades
• Louis Iverson, Anantha M. Prasad, Stephen Matthews, Matthew Peters. 2014. Climate as an agent of change in forest landscapes
• Matthew Peters, Louis Iverson, Stephen Matthews. 2014. Spatio-temporal trends of drought by forest type in the conterminous United States, 1960-2013
• Maria Janowiak, Chris Swanston, Linda M. Nagel, Leslie Brandt, Patricia R. Butler, Stephen Handler, P. Danielle Shannon, Louis Iverson, Stephen Matthews, Anantha M. Prasad, Matthew Peters. 2014. A practical approach for translating climate change adaptation principles into forest management actions
• Maria Janowiak, Louis Iverson, David J. Mladenoff, Emily Peters, Kirk R. Wythers, Weimin Xi, Leslie Brandt, Patricia R. Butler, Stephen Handler, P. Danielle Shannon, Chris Swanston, Linda R. Parker, Amy J. Amman, Brian Bogaczyk, Christine Handler, Ellen Lesch, Peter B. Reich, Stephen Matthews, Matthew Peters, Anantha M. Prasad, Sami Khanal, Feng Liu, Tara Bal, Dustin Bronson, Andrew Burton, Jim Ferris, Jon Fosgitt, Shawn Hagan, Erin Johnston, Evan Kane, Colleen Matula, Ryan O'Connor, Dale Higgins, Matt St. Pierre, Jad Daley, Mae Davenport, Marla R. Emery, David Fehringer, Christopher L. Hoving, Gary Johnson, David Neitzel, Michael Notaro, Adena Rissman, Chadwick Rittenhouse, Robert Ziel. 2014. Forest ecosystem vulnerability assessment and synthesis for northern Wisconsin and western Upper Michigan: a report from the Northwoods Climate Change Response Framework project
• Stephen Handler, Matthew Duveneck, Louis Iverson, Emily Peters, Robert M. Scheller, Kirk R. Wythers, Leslie Brandt, Patricia Butler, Maria Janowiak, P. Danielle Shannon, Chris Swanston, Kelly Barrett, Randy Kolka, Casey McQuiston, Brian J. J. Palik, Peter B. Reich, Clarence Turner, Mark White, Cheryl Adams, Anthony D'Amato, Suzanne Hagell, Patricia Johnson, Rosemary Johnson, Mike Larson, Stephen Matthews, Rebecca Montgomery, Steve Olson, Matthew Peters, Anantha M. Prasad, Jack Rajala, Jad Daley, Mae Davenport, Marla R. Emery, David Fehringer, Christopher L. Hoving, Gary Johnson, Lucinda Johnson, David Neitzel, Adena Rissman, Chadwick Rittenhouse, Robert Ziel. 2014. Minnesota forest ecosystem vulnerability assessment and synthesis: a report from the Northwoods Climate Change Response Framework project
• Dennis S. Ojima, Louis Iverson, Brent L. Sohngen, James Vose, Christopher W. W. Woodall, Grant M. Domke, David L. Peterson, Jeremy S. Littell, Stephen Matthews, Anantha M. Prasad, Matthew Peters, Gary W. Yohe, Megan M. Friggens. 2014. Risk assessment [Chapter 9]
• Stephen Matthews, Louis Iverson, Matthew Peters, Anantha M. Prasad, Sakthi Subburayalu. 2014. Assessing and comparing risk to climate changes among forested locations: implications for ecosystem services
• Stephen Handler, Matthew Duveneck, Louis Iverson, Emily Peters, Robert M. Scheller, Kirk R. Wythers, Leslie Brandt, Patricia Butler, Maria Janowiak, P. Danielle Shannon, Chris Swanston, Amy Clark Eagle, Joshua G. Cohen, Rich Corner, Peter B. Reich, Tim Baker, Sophan Chhin, Eric Clark, David Fehringer, Jon Fosgitt, James Gries, Christine Hall, Kimberly R. Hall, Robert Heyd, Christopher L. Hoving, Ines Ibáñez, Don Kuhr, Stephen Matthews, Jennifer Muladore, Knute Nadelhoffer, David Neumann, Matthew Peters, Anantha M. Prasad, Matt Sands, Randy Swaty, Leiloni Wonch, Jad Daley, Mae Davenport, Marla R. Emery, Gary Johnson, Lucinda Johnson, David Neitzel, Adena Rissman, Chadwick Rittenhouse, Robert Ziel. 2014. Michigan forest ecosystem vulnerability assessment and synthesis: a report from the Northwoods Climate Change Response Framework project
• Leslie Brandt, Hong He, Louis Iverson, Frank R. Thompson, Patricia Butler, Stephen Handler, Maria Janowiak, P. Danielle Shannon, Chris Swanston, Matthew Albrecht, Richard Blume-Weaver, Paul Deizman, John DePuy, William D. Dijak, Gary Dinkel, Songlin Fei, D. Todd Jones-Farrand, Michael Leahy, Stephen Matthews, Paul Nelson, Brad Oberle, Judi Perez, Matthew Peters, Anantha M. Prasad, Jeffrey E. Schneiderman, John Shuey, Adam B. Smith, Charles Studyvin, John M. Tirpak, Jeffery W. Walk, Wen J. Wang, Laura Watts, Dale Weigel, Steve Westin. 2014. Central Hardwoods ecosystem vulnerability assessment and synthesis: a report from the Central Hardwoods Climate Change Response Framework project
• Matthew Peters, Stephen Matthews, Louis Iverson, Anantha M. Prasad. 2013. Delineating generalized species boundaries from species distribution data and a species distribution model
• Matthew Peters, Louis Iverson, Anantha M. Prasad, Stephen Matthews. 2013. Integrating fine-scale soil data into species distribution models: preparing Soil Survey Geographic (SSURGO) data from multiple counties
• Louis Iverson, Matthew Peters, Stephen Matthews, Anantha Prasad. 2013. An overview of some concepts, potentials, issues, and realities of assisted migration for climate change adaptation in forests
• Daniel Yaussy, Louis Iverson, Stephen Matthews. 2013. Competition and climate affects US hardwood-forest tree mortality
• Matthew Peters, Louis Iverson, Stephen Matthews, Anantha M. Prasad. 2013. Wildfire hazard mapping: exploring site conditions in eastern US wildland—ruban interfaces
• Anantha M. Prasad, Judith D. Gardiner, Louis Iverson, Stephen Matthews, Matthew Peters. 2013. Exploring tree species colonization potentials using a spatially explicit simulation model: implications for four oaks under climate change
• Megan M. Friggens, Stephen Matthews. 2012. Risk assessment for two bird species in northern Wisconsin
• Louis Iverson, Stephen Matthews, Anantha M. Prasad, Matthew Peters, Gary W. Yohe. 2012. Appendix 2: Risk-based framework and risk case studies. Risk assessment for forested habitats in northern Wisconsin.
• Megan M. Friggens, Stephen Matthews. 2012. Appendix 2: Risk-based framework and risk case studies. Risk Assessment for two bird species in northern Wisconsin.
• Louis Iverson, Stephen Matthews, Anantha M. Prasad, Matthew Peters, Gary Yohe. 2012. Development of risk matrices for evaluating climatic change responses of forested habitats
• Anantha M. Prasad, Louis Iverson, Matthew Peters, Stephen Matthews. 2011. Using a cellular model to explore human-facilitated spread of risk of EAB in Minnesota
• Louis Iverson, Anantha M. Prasad, Kathleen Knight, Daniel A. Herms, Stephen Matthews, Matthew Peters, Annemarie Smith, Robert P. Long. 2011. Potential replacements for northwoods black ash in a changing climate: the confluence of two challenges
• Stephen Matthews, Louis Iverson, Anantha M. Prasad, Matthew Peters. 2011. Changes in potential habitat of 147 North American breeding bird species in response to redistribution of trees and climate following predicted climate change
• Louis Iverson, Anantha M. Prasad, Stephen Matthews, Matthew Peters. 2011. Lessons learned while integrating habitat, dispersal, disturbance, and life-history traits into species habitat models under climate change
• Stephen Matthews, Louis Iverson, Anantha M. Prasad, Matthew Peters, Paul G. Rodewald. 2011. Modifying climate change habitat models using tree species-specific assessments of model uncertainty and life history-factors
• Chris Swanston, Maria Janowiak, Louis Iverson, Linda Parker, David Mladenoff, Leslie Brandt, Patricia Butler, Matt St. Pierre, Anantha M. Prasad, Stephen Matthews, Matthew Peters, Dale Higgins, Avery Dorland. 2011. Ecosystem vulnerability assessment and synthesis: a report from the Climate Change Response Framework Project in northern Wisconsin
• Anantha M. Prasad, Louis Iverson, Stephen Matthews, Matthew Peters. 2010. Climate change poses additional threat to the future of ash resources in the eastern United States
• Louis Iverson, Anantha M. Prasad, Stephen Matthews, Matthew Peters. 2010. Merger of three modeling approaches to assess potential effects of climate change on trees in the eastern United States
• Stephen Matthews, Louis Iverson, Anantha M. Prasad, Matthew Peters. 2010. Potential impacts of climate change on bird and tree habitats within the Appalachian Mountains
• Jessica J. Hellmann, Knute J. Nadelhoffer, Louis Iverson, Lewis H. Ziska, Stephen Matthews, Philip Myers, Anantha M. Prasad, Matthew Peters. 2010. Climate change impacts on terrestrial ecosystems in metropolitan Chicago and its surrounding, multi-state region
• Anantha M. Prasad, Louis Iverson, Matthew Peters, Jonathan M. Bossenbroek, Stephen Matthews, T. Davis Sydnor, Mark W. Schwartz. 2010. Modeling the invasive emerald ash borer risk of spread using a spatially explicit cellular model
• Louis Iverson, Anantha M. Prasad, Stephen Matthews, Matthew Peters, Coeli Hoover. 2010. Potential changes in habitat suitability under climate change: lessons learned from 15 years of species modelling
• Louis Iverson, Anantha M. Prasad, Stephen Matthews, Matthew Peters. 2009. Potential changes in tree habitat under Illinois climate change. Chapter 16
• Anantha M. Prasad, Louis Iverson, Stephen Matthews, Matthew Peters. 2009. Atlases of tree and bird species habitats for current and future climates
• Louis Iverson, Anantha M. Prasad, Stephen Matthews, Matthew Peters. 2008. Estimating potential habitat for 134 eastern US tree species under six climate scenarios
• N.L. Rodenhouse, Stephen Matthews, K.P. McFarland, J.D. Lambert, Louis Iverson, Anantha M. Prasad, T.S. Stillett, R.T. Holmes. 2008. Potential effects of climate change on birds of the Northeast
• Louis Iverson, Anantha M. Prasad, Stephen Matthews. 2008. Modeling potential climate change impacts on the trees of the northeastern United States
• Louis Iverson, Anantha M. Prasad, Stephen Matthews, Matthew Peters. 2007. Potential impacts of climate change on birds and trees of the eastern United States: newest climate scenarios and species abundance modelling techniques
• Mark W. Schwartz, Louis Iverson, Anantha M. Prasad, Stephen Matthews, Raymond J. O'Connor, Raymond J. O'Connor. 2006. Predicting extinctions as a result of climate change
• Louis Iverson, Anantha M. Prasad, Mark Schwartz, Stephen Matthews. 2006. Statistical modeling of current and potential future habitat, then simulation of potential colonization within that habitat, for tree species in the Eastern US
• Stephen Matthews, Raymond O'Connor, Louis Iverson, Anantha M. Prasad. 2004. Atlas of climate change effects in 150 bird species of the Eastern United States

Data and Tools

• Tool
  

  Climate Change Tree Atlas

  https://research.fs.usda.gov/nrs/products/dataandtools/climate-change-tree-atlas
• Tool
  

  Climate Change Bird Atlas

  https://research.fs.usda.gov/nrs/products/dataandtools/climate-change-bird-atlas

• Program
  

  Climate and Carbon Cycle Sciences

  https://research.fs.usda.gov/nrs/programs/climate
  Station

  Northern Research Station
• Project
  

  Species Habitat Modeling Under Future Climates

  https://research.fs.usda.gov/nrs/projects/species-habitat-modeling
  Station

  Northern Research Station
• Project
  

  Species Habitat Modeling Under Future Climates

  https://research.fs.usda.gov/nrs/projects/habitatmodeling
  Station

  Northern Research Station

Multimedia

• Webinars

  Climate Change Tree Atlas: New data and new opportunities in version 4

  https://research.fs.usda.gov/nrs/products/multimedia/webinars/climate-change-tree-atlas-new-data-and-new-opportunities-version-4

• Ohio State University, Columbus, Ohio, Ph.D., Natural Resources, 2008
• University of Maine, Orono, Maine, M.S., Wildlife Ecology, 2003
• Frostburg State University, Frostburg, Maryland, B.S., Wildlife Biology, 1997